B. Papp szerk.: Studia Botanica Hungarica 30-31. 1999-2000 (Budapest, 2000)

Erdei, Boglárka; Lesiak, Maria: A study of dispersed cuticles, fossil seeds and cones from Sarmatian (Upper Miocene) deposits of Sopron-Piusz puszta (W Hungary)

22.5-37.5 |jm. Anticlinal cell walls are strongly thickened. Stomata are randomly arranged and their shape is more circular than that of the other specimens. The guard cells and stomatal pores with their 48-53 urn and 18.75-37.5 urn length are considerably large. Type of stomata is cyclocytic with one subsidiary cell on one pole and two subsidiaries on the other. Guard cells are sunken. Outer stomatal ledge is strongly thickened and sometimes weakly developed T-shaped thickenings are noticeable. Taxodiaceae gen. et sp. 2 (Figs 8-9) Material: BP 99.497.1 ; BP 99.498.1 ; BP 99.499.1. Description: Fragments of cuticles are of medium thickness. Due to poor preservation the anticlinal cell walls are hardly observable. Stomata are more elongated and their arrangement seems to be more regular than that of the specimen No. BP 99.496.1 of Taxodiaceae gen. et sp. 1, but not parallel. The length of guard cells (outer stomatal ledge) is 47-52 urn, their width is 31.3-37.5 um. The length of stomatal pores in specimens No. BP 99.497.1 and BP 99.499.1 are 12.5-21.3 urn and 20-25 urn, respectively. The latter one seems to have larger stomatal pores, however, it should be taken into consideration that the latter fragment must have been strongly compressed, since the stomata are also narrower and more elongated. Type of stomata is cyclocytic with sunken guard cells. Well developed, T-shaped thickenings are striking. Discussion: Stomata of Taxodium dubium (Sternb.) Heer described from the Pliocene flora of Ruszów by HUMMEL (1983, p. 15, PI. 2, Figs 1-6) are well comparable. According to the description made by HUMMEL, leaves are amphistomatic, orientation of stomata is variable but generally their longitudinal axis is not parallel with that of the leaf. Type of stomata is amphi- or monocyclocytic. Stomata are sunken and the outer stomatal ledge forms a thickened annulus on the surface of the epidermis. T-shaped thickenings are often developed but not generally present (HUMMEL'

s opinion is that the presence of T-shaped thickenings depends also on the degree of maceration). FERGUSON (1971) described some Taxodium remains (PI. 2, Figs E-F) from the Miocene flora of Kreuzau and remarked that species of Taxodiaceae provide very similar characteristic features of the cuticle, however, that of Taxodium and Glyptostrobus are distinguishable. Stomata of Glyptostrobus are mostly arranged parallel to the longitudinal axis of the leaf, but those of Taxodium are either aligned transversely to the length of the leaf or orientated in various directions. Unfortunately, the cuticle