L. Lőkös szerk.: Studia Botanica Hungarica 24. 1993 (Budapest, 1993)

Hably, Lilla: Egerian macroflora from the Andornaktálya sandpit (Hungary)

1959 Anacardiíes cfr. Schinus molle L.; Andreánszky, p. 151, Text-Fig. 174, Pl. 45, Fig. 5. 1959 Myrica lignitum (Unger) Saporta; Rásky, p. 454. 1963 Schinus oligocaenicum Andreánszky et Nóvák; Andreánszky, p. 100, Pl. 2, Fig. 13. 1976 Engelhardtia détecta Saporta; Knobloch and Kvacek, p. 27, Pl. 10, Figs 2-7; Pl. 11, Fig. 3; Pl. 12, Figs 1, 2, 8; Pl. 17, Fig. 12; Pl. 19, Fig. 6; Pl. 20, Fig. 2. 1977 Engelhardtia orsbergensis (Wessel et Weber) Jähnichen, Mai et Walther; Jähnichen et al, p. 323, PI. 9, Fig. 4. 1981 Engelhardtia orsbergensis (Wessel et Weber) Jähnichen, Mai et Walther; Pálfalvy, p. 492. 1984 Palaeocarya orsbergensis (Wessel et Weber) Jähnichen, Friedrich et Takác; Jähnichen et al, p. 110, Pl. 1, Figs 1-6; Pl. 2, Figs 1, 2; Pl. 3, Figs 1, 3; Pl. 4, Figs 1-6; Pl. 5, Figs 1-7. 1985 Engelhardtia orsbergensis (Wessel et Weber) Jähnichen, Mai et Walther; Hably, p. 106, Pl. 20, Fig. 5; Pl. 24, Figs 3-6; Pl. 25, Figs 1-8; Pl. 26, Figs 1-6; Pl. 27, Fig. 5. Material: upper: 89.556.1.; cf. 89.561.1.; 89.609.1. Description: Two, almost complete specimens, 9.1 cm and 11.5 cm long, 1.0 and 1.5 cm wide, respectively. Shape lanceolate, apex acute, rounded, base asymmetrical. Margin toothed, small, barbed teeth, arched upwards. Very widespread species in the European Tertiary as well as in Hungary. It ranges from the Eocene to Late Miocene here. There are mass occurrences in the Tard Clay Formation and at . Ipolytarnóc, and significant localities in the Egerian, occurring in masses in a few localities; otherwise, there are at least a few specimens everywhere. It is a thermophylous palaeotropical element. Mostly associated with Daphnogene, Laurus, Laurophyllum, and Platanus neptuni. "Rhamnus" warthae Heer 1872 Rhamnus warthae Heer; Heer, p. 23, Pl. 5, Figs 2, 3; Pl. 6, Figs 3, 4, 5. 1872 Juglans heerii Ettingshausen; Heer, p. 24, Pl. 5, Fig. 4a. 1887 Rhamnus warthae Heer; Staub, p. 360, Pl. 38, Figs 1, 2. 1887 Juglans heerii Ettingshausen; Staub, p. 281, Pl. 27, Fig. 1. 1887 Laurusprimigenia Unger; Staub, p. 303, PI. 28, Fig. 6; PI. 29, Fig. 1; PI. 34-35, Fig. Id. Material: 89.557.2.; cf. 89.564.1. Description: An almost complete, measurable specimen, 9.8 cm long, 3.1 cm wide. Shape el­liptic, venation camptodromous, there are rich loop networks at the terminations of secondary veins. The margin is damaged, with very rare and small teeth. Many imprints have been found in the Wind-brickyard named by ANDREÁNSZKY (1966) as Symplocos and Elaeocarpus palaeolanceolatus . STAUB (1887) published the species under the names Laurus primigenia, Juglans heerii, J. ungeri, J. bilinica from the Upper Oligocène Zsil (Jiu) valley flora (Romania). New Egerian localities (Kesztölc, Nagysáp) also yielded the species, described as Symlocos sp. (HABLY 1988, 1989). It is a significant accessory element at Pomáz. The species is restricted to the inner Carpathian region. Known localities are in the Upper Oligocène of Hungary and Transylvania. Since there are no localities of the same age in Yugoslavia and Bulgaria, farther connections cannot be traced. The well-known, published flo­ras of the north (Bohemia, Germany) do not contain this species, not even as a synonym. We suggest, that "Rhamnus" warthae was an endemic species of the inner Carpathian region, ap­pearing in the Upper Oligocène. (Its taxonomic position is unknown; Rhamnus is preserved for priority's sake.) It clearly belongs to a family of a thermophylous tropical-subtropical family. 12

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