Zs. P. Komáromy szerk.: Studia Botanica Hungarica 15. 1981 (Budapest, 1981)

Dobolyi, Konstantin, Szabó, László , Szerdahelyi, Tibor; Szujkó-Lacza, Júlia: Data to the Genisto pilosae-Quercetum and the flora of the Bükk Mountains

3, 4), we believe that the presence of these oak-forests depend on the pH value of the forest soil developed on acidiferous fundamental rock. There is an old granite block in Velence Mts. and a small part of acidic gritstone in Buda Mts. on which the Querceto-Luzuletum subcarpaticum had developed. The Genisto pilosae-Quercetum occurs in schist stone in the Bükk Mts. The Genista pilosa, Luzula alblda, Veronica officinalis. Viscaria vulgaris belong to the relatively acidiferous group of plants in Hungary (CSAPODY, HORANSZKY, PÓCS, SIMON, SZODFRTDT, TALLÓS in: MAYER 1962). In both localities Avenella flexuosa and Vaccinium myrtillus occur, too. They indicate the low pH value in the soil (Table 1). Since the size of samples, the recording time of the samples are different, we have analysed only the samples in Odorhegy in detail. The most frequent species are the next: F = 10 Quercus petraea, Avenella flexuosa, Luzula albida, Melampyrum cristatum; F = 7 Vaccinium myrtillus; F = 6 Genista piloBa. Veronica offici nalis; F = 5 Pinus silvestris cult., Brachypodium silvaticum in Odorhegy. Based on these results the acidiferous oak-forest is believed to be extrazonal in Hungary. Geographically, and on the basis of species composition the localities of Genisto pilosae-Quercetum in the Bükk MtB. may be near the zonal acidiferous oak-forest. In decreasing order of the H'-^ value of the more frequent species in Table 3: Quercus pet raea

, Avenella flexuosa

, Melampyrum cristatum. Genista pilosa, Veronica officinalis, Vaccinium myrtillus, Luzula albida. The other species are less frequent in samples and they have much lower H'JJ values. Values of the evenness lighten the species in a more realistic form than the diversity values. The H*R values of the Individual species are more determined by the frequency than the differences among sizes of patches In a row in Table 3. (It is a well-known character of the SHANNON'S information model.) The number of species and their covering values are different in each per samples (Table 2.) The number of species is extremely high in sample No. 5, which is transitional and situated near the bottom of the valley. The H*Q values of the samples are decreasing as follows: No. 5, 3, 1, 4, 2, 8, 7, 6 and 10. The values of evenness do not follow this order of samples. Comparing the Bize of patches, diversity values in H'

R amd H'Q we can conclude that some of the previous species may show higher number than the latter in the Genisto pilosae-Quercetum here. We suppose that the covering behaviour of any species in ten samples also may give some information of this character of the individual species - H' value - refer to the area of it as a whole. Here those species produce a higher diversity value, which expand mainly or totally with vegetative (Table 3). They occur more frequently also in thiB community. It is a well-known fact that the species composition within one sample or community depends on numerous factors. In this case we can suppose that one of these factors is the manner of the species* expand and as a result of this, the size of patches. The size of patches of species are interdependent on each other but the nature and comprise of these relations are hardly known. Saturation of samples or community The surface of soil is almost completely covered with the two layers (i.e. trees and herbs) in the studied samples of Genisto pilosae-Quercetum. So saturation by different vascular plants is more than 100 per cent with respect to the covering value (Table 2). But the number of species in the same quadrates is low and strictly limited (8-14) except in the sample No. 5. Based on these facts the question arises, whether they are characteristic for the Genisto pilosae-Quercetum , being hardly saturated by the species (compared with other communities) but well or over saturated by the size of different species' populations. It is seems to be true. Following the comparison, the H' and J values refer to the H'

C and J

c

: H*rj and JD: also the H'-^ and Jjj. The real saturation of soil is caused much more by the flowering and cryptogamous species but the latters are present often only In spores or myceliums. Consequently, we can account on only those species which are well determinable during the vegetation period.