Zs. P. Komáromy szerk.: Studia Botanica Hungarica 12. 1977 (Budapest, 1977)

Radics, Ferenc: The identification of Rorippa species and hybrids (Cruciferae) based on external morphological features of their seeds

repens BORB. (R. amphibia x silvestris), R, prostrata (BERG. ) SCHINZ and THELL (R. amphibia x islandica ? ). In the same book, he mentioned also 10 further hybrids, occurring spora­dically or to be recognized with difficulty, e. g. R. neogradensis BORB. (R. islandica x austriaca), R. menyhártiana BORB. (R. islandica x sil­vestris), R. küllődensis PROD. (R. kerneri x amphibia). Recently SOO (1968) discriminated in his synopsis altogether 112 taxa due regard to the territory of present-day Hungary, taking into account the above systématisations and with the results of researches from abroad. He begins the description of the Rorippa genus with the following words: "This genus is the most difficult one among the Cruciferaet it is a com­plex of species, hybridogen stabilized species, transition forms and hy­brids of species. Origin and value of various taxa can be cleared up later only, by genetic examinations and analyses of population. " (1. c. 359). SOO described four basic species in detail: R islandica (palustris resp. JONSELL 1968), R. austriaca , R. amphibia and R. silvestris ; and one subspecies: R. silvestris ssp. kerneri and three hybrids: Rorippa x pro­strata (BERG. 1786 sub Myagro) SCH. et TH. 1913; Rorippa x astylis (RCHB. 1932 sub Nasturtio) RCHB. 1838 = syn. R. barbaraeoides (TAUSCH 1840 sub Nasturtio) CE LAK, 1874 and R. armoracioides TAUSCH (1840 sub Nasturtio) FUSS 1866. The second hybrid has 3 sub­species: R x astylis ssp. astylis (barbaraeoides var. astyla typica NYAR 1955), R. x astylis ssp. reichenbachii (KNAF in OPIZ 1852 sub Nastur­tio) SOÓ 1966 and R. x astylis ssp. capillipes (BORB. 1379 p. sp. ) SOO 1966. As to this latter, he says that it is -the hybrid R. austriaca x sil­vestris (1. c. 365). Two subspecies belong to the third hybrid: R. x armo­racioides (var. pinnatifidum TAUSCH) and R. x armoracioides ssp. ter­restris (TAUSCH 1840 sub Nasturtio p. sp. ) JAV, 1924. According to SOO, R. x prostrata is probably a stabilized hybrid of R. silvestris and R. amphibia . R. astylis is originated from R. islandica x R. silvestris . and R. x armoracioides from R. austriaca and R, silvestris As to this latter hybrid, he refers to FRÖHLICH (1914). Further on, he enumerates 9 hybrids saying that the origin of hybrids is "partly dubious". Moreover, he enumerates 8 hybrids (I.e. 367). Lately, a revision made by TOMSOVIC ( 1969) stated the occurrence of 5 basic species in the ter­ritory of present-day Czechoslovakia, thus partly in the Carpathian Basin: R. amphibia , R. palustris , R. austriaca , R. silvestris and R. pyrenaica , further 3 hybrids: R. x hungari ca BORB. 1877 (R. amphibia x austriaca), R. x barbaraeoides (TAUSCH) (5ELAK. 1874 (= R. amphibia x silvestris) and the R. x armoracioides (TAUSCH) FUSS 1866 (= R. austriaca x sil­vestris). He also inserted H. x prostrata (BERG. ) SCH. et TH. into the sphere of R. x barbaraeoides (TAUSCH) £ELAK and R. terrestris <5ELAK, into that of R. x armoracioides . Formerly, BORBAS (1878) thought that the taxon

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