Dr. Murai Éva szerk.: Parasitologia Hungarica 19. (Budapest, 1986)

Trianchoratus pahangensis sp.n. (Fig. 57-62). Host: ppWcejjhalus Jucius_(Cuvier et Vallenciennes) (Ophicephalidae)Locality: Tasek Bera, Pahang. Type specimens: Holotype USNM No. 78040, paratype UMZD (P) No. 27. No. measured: 11. Body size 477 (336-714) x 118 (84-168). 4 eyespots, posterior pair slightly larger. Alimentary system as in T. malayensis sp. n. Ventral lobes more developed and flexible than dorsal lobes. Dorsal anchors unequal, right anchor rudimentary, somewhat arcuate, length 16(16-18), left anchor developed, inner length 37 (35-40), outer length 25 (2 5-30), inner root 22 (20-2 5), outer root 8 (5-10) and recurved point 15 (12-15). Ventral anchors equally developed, inner length 35 (35-40), outer length 26 (25-28), inner root 18 (16-20), outer root 6(5-9) and recurved point 16(15-18). All developed anchors oriented ventrally. Haptoral bar absent. 14 marginal hooks, proximal tip with knob resembling a pin-head, length 21 (20-24). Copulatory organ consisting of slightly coiled tube and grooved accessory piece. Vagina dextral with sclerotised funnel-like opening and short vaginal tube. Remarks This species differs from T. malayensis sp.n. in the structures of the developed anchors. In this species the anchors are almost similar in size and shape but not in T. malayensis where they are structurally dissimilar. This species is named Trianchoratus pahangensis sp. n. after the state of Pahang. Trianchoratus ophicephali sp.n. (Fig. 50-56). Host: Ophicephalus striatus (Bloch) (Ophicephalidae). Locality: Tasek Bera, Pahang (type locality), Bukit Merah Reservoir, Perak. Type specimens: Holotype USNM No. 78039, Paratypes in USNM No. 77284 and UMZD(P) No. 26. No. measured: 15. Body size 767 (532-1013) x 215 (138-288). 4 eyespots. Alimentary system and reproductive systems as in T. malayensis sp.n. (see Fig. 12). Dorsal anchors unequal, right anchor rudimentary, somewhat arcuate, length 22 (20-25); left dorsal anchor developed, inner length 36 (35-40), outer length 22 (20-25), inner root 18 (18-20), outer root 8 (5-10) and recurved point 20 (20-22). Ventral anchor equally developed, inner length 50 (49-50), outer length 26 (25-30), inner root 32 (30-35), outer root 5 (5-8) and recurved point 18 (20-25). Haptoral bar absent. 14 marginal hooks, proximal tip with knob resembling pin-head, length 20 (20-24). Copulatory organ consists of slightly coiled tube and grooved accessory piece. 2 prostate glands. Vagina dextral with sclerotised funnel-like opening and short vaginal tube. Remarks The present species differs from T. malayensis sp.n. in having almost equal sized anchors compared to T. malayensis where the ventral anchors are larger. This species is different from T. pahangensis sp.n. in having shorter points. This species is named Trianchoratus ophicephali sp. n. after the host species. DISCUSSION The position of the anchors have been elucidated by studying live specimens. The single well developed anchor is located on the dorsal side, while the 2 well developed anchors are located ventrally on separate lobes, and all the anchors are oriented ventrally. The larval forms of Trianchoratus species from 0._lucius indicate that the paired anchors complete their development before the single well developed anchor. According to GUSSEV (1978) the ventral anchors usually developed first before the dorsal anchors in four anchor monogeneans. Hence if Trianchoratus follows the same trend in development then the single well developed anchor