Matskási István (szerk.): A Magyar Természettudományi Múzeum évkönyve 102. (Budapest 2010)
Kirschenhofer, E.: New and little-known species of Carabidae from the Middle East and Southeast Asia Coleoptera, Carabidae: Lebiini, Brachinini)
New and little-known species of Car abidaefrom Asia (Coleoptera, Carbidae) 31 Coptodera johorensis sp. n. (Fig. 22) Type material - Holotype (female): "Malaysia - W, Johor, 15 km NM of Kota Tinggi, Muntahak mt„ 200 m, 7.-13.III.2002, P. Cechovsky" (CAD). Description - Habitus: Fig. 22. Length 8.0 mm, width 3.7 mm. Colour and lustre: head reddish brown, from posterior margin of clypeus to base with a rather wide median black band which continues over the whole pronotal midline (from anterior margin to base), remainder of pronotum reddish brown; elytra black; marginal groove reddish yellow; elytra with two reddish yellow spots, anterior covering intervals 3 to 8, slightly abbreviated on 3, posterior covering intervals 2 to 4, on 2 and 3 shortened anteriorly, on 3 and 4 posteriorly slightly extended; clypeus darkened, mandibles reddish brown with outer margin narrowly darker; clypeus, palpi and tarsi reddish yellow; antennal segments dark reddish yellow, 1 and 3 darker; femora and tibiae darker, with protibia lighter reddish on insides; dorsum moderately shiny, glabrous; underside reddish yellow, elytral epipleura brownish, anterior margins of segments darker; underside shiny, glabrous. Head: with large protruding eyes; frons convex; temples short, weakly developed; frontal depressions broad, shallow; labrum with a short furrow-like median depression behind anterior margin; head without distinct diagnostic features. Pronotum: 1.93 times as wide as long, weakly convex, with sides roundely narrowing anteriorly, widest before middle, posterior angles sharp, slightly obtuse, sides towards these weakly and obliquely narrowing; marginal groove wide, distinctly widening towards base; anterior margin straight; posterior margin arcuate in middle, base bilaterally weakly declivous; basal impressions with widened pits; median line very finely incised, hardly reaching anterior margin or base. Elytra: broadly oval, weakly convex; basal margin complete, roundly connected to lateral margin; humeri distinct, not declivous; marginal groove wide; outer angles of apical incision shortly rounded, apex of elytral suture shortly rounded; striae deep, very finely punctate at bottom; intervals smooth, convex, third with 3 setigerous punctures of which anterior adjacent to third stria just behind anterior margin, posterior adjacent to second stria. Microsculpture: head with minute polygons, pronotum indistinctly transversely reticulate, elytra with more densely transverse meshes. Distribution - Malaysia. Etymology - It is named after the place of origin. Comparisons - The new species resembles Coptodera tetrastigma CHAUDOIR, 1869 (type locality: "Borneo, Sarawak"). It is, however, larger than C. tetrastigma (6.0 mm) and with spots rather evenly rounded (spots larger and posterior one "U"-shaped in the middle and serrated posteriorly in C. tetrastigma). The head and pronotum of C. tetrastigma lack the dark median band. Annls hist.-nat. Mus. natn. hung. 102, 2010