Matskási István (szerk.): A Magyar Természettudományi Múzeum évkönyve 98. (Budapest 2006)

Makranczy, Gy.: Systematics and phylogenetic relationships of the genera in the Carpelimus group (Coleoptera: Staphylinidae: Oxytelinae)

ous as its placement is weakly supported (no unique synapomorphies). The primary result of this analysis, is that all the characters used for Oxytelini as recognized earlier either (1) vary within the tribe or (2) occur scattered in other genera outside the tribe. I am also guessing that the larval characters will confirm this conclusion, as well, but they definitely require further study. As the analysis concentrates primarily on the group of genera that was previously considered to be a monophyletic group within the former Thinobiini, the taxon sampling is limited for the Bledius, Thinobius, Oxytelus and Anotylus groups of genera. Therefore relationships among these taxa cannot be reliably inferred from the phylogenetic hypothesis depicted on the trees presented (Figs 2-3). The sampling of these taxa was only aimed at inclusion of characters and character states that might give information on the relatedness of the ingroups. This refers to the relationships of Neoxus and Thinobius, the genera of the former Oxytelini among each other, the species groups within Bledius (it was discussed in HERMAN 1986) and the basal clades of Deleasterini and Coprophilini. The sizes and diversities of some of these taxa is such, that their adequate sampling was impossible within the present study. SCHEERPELTZ in 1944 erected a new tribe, Trigonobregmini, and described the genus Trigonobregma, based on the species Thinobius ocularis FAUVEL, 1902. HERMAN (2001) listed the genus and species in Oxytelinae and in a further paper (HERMAN 2003) explained that it was overlooked at the time of his 1970 generic revision and he erroneously included T. ocularis in Thinobius. Upon examination of the single syntype specimen it is obvious that it is not a member of the Oxytelinae, because there are not seven visible abdominal sternites, not even a trace of a developed second sternite. The species belongs in the tribe Aphaenostemmini (subfamily Omaliinae), but transferring the name is outside the scope of this work, so will be dealt with elsewhere. The situation with the Oxytelus and Anotylus groups of genera (although only represented by 3 species in the analysis, was studied extensively on a worldwide basis) is dealt with in a separate chapter. The characters used in the analysis of the Carpelimus group do not provide resolution for these clades - in fact, probably no known morphological characters would. THE PROPOSED NEW TRIBAL CLASSIFICATION 1. Tribe Deleasterini REITTER, 1909 (type: Deleaster) = Pholidiens MULSANT et REY, 1877 (type: Pholidus) = Pholidini ACLOQUE, 1896 (type: Pholidus) = Euphaniae REITTER, 1909 (type: Euphanias)