Matskási István (szerk.): A Magyar Természettudományi Múzeum évkönyve 98. (Budapest 2006)

Makranczy, Gy.: Systematics and phylogenetic relationships of the genera in the Carpelimus group (Coleoptera: Staphylinidae: Oxytelinae)

Anotylus THOMSON, 1859 = Oxytelopsis FAUVEL, 1895, syn. n. = Rimba BLACKWELDER, 1952, syn. n. = Styloxys DES GOZIS, 1886 = Oxytelodes BERNHAUER, 1908 = Emopotylus BERNHAUER, 1910 = Boettcherinus BERNHAUER, 1936 = Oncoparia BERNHAUER, 1936 = Paracaccoporus STEEL, 1948 = Oxytelosus CAMERON, 1950 = Microxytelus FAGEL, 1956 = Oxytelops FAGEL, 1956 = Pseudodelopsis FAGEL, 1957 = Anotylops FAGEL, 1957 Characters: crest shaped scutellar impression; basolateral ridge missing on tergite II. PHYLOGENETIC ANALYSIS Introduction and methods In phylogenetic analyses that are made on large and very diverse groups, there can be two basic problems: the groups (and the species that represent them in the analysis) are not those that can answer questions; and the characters selected are not those that carry information on relatedness. Of course, the larger the group, the harder it is to avoid these difficulties. It is simply not possible for a specialist to dis­sect and study every single specimen in those specious groups and gain a complete understanding of variability and the potential phylogenetic value of characters. The data matrix (Table 1) was created in WINCLADA and was analyzed with the parsimony ratchet as implemented in NONA. With multistate characters consid­ered nonadditive (Fitch parsimony), 32 most parsimonious trees are recovered (L=258, CI=37, RI=65). One of them is presented on Fig. 3. Bootstrap values and Bremer support for the branches are displayed on this tree. Bremer support values were calculated in NONA using the commands hold 15000, suboptimal 20, bsupport 10. Bootstrap values were calculated using NONA executed from Winclada, using 1000 replications.

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