Matskási István (szerk.): A Magyar Természettudományi Múzeum évkönyve 98. (Budapest 2006)
Makranczy, Gy.: Systematics and phylogenetic relationships of the genera in the Carpelimus group (Coleoptera: Staphylinidae: Oxytelinae)
genus Thinodromus KRAATZ, 1857 (= Apocellagria CAMERON, 1920, syn. n.) genus Trogactus SHARP, 1887 genus Thinobius KIESEN WETTER , 1844 genus Neoxus HERMAN, 1970 genus Sciotrogus SHARP, 1887 4. Tribe Oxytelini THOMSON, 1858 genus Sartallus SHARP, 1871 genus Parosus SHARP, 1887 genus Paraploderus HERMAN, 1970 genus Ecitoclimax BORGMEIER, 1934 genus Jerozenia HERMAN, 2004 genus Platystethus MANNERHEIM, 1830 genus Oxytelus GRAVENHORST, 1802 (= Paroxytelopsis CAMERON, 1933, syn. n., = Anisopsidius FAGEL, 1960, syn. n., = Anisopsis FAUVEL, 1904, syn. n., = Hoplitodes FAUVEL, 1904, syn. n.) genus Apocellus ERICHSON, 1839 genus Anotylus THOMSON, 1859 (= Rimba BLACKWELDER, 1952, syn. n., = Oxytelopsis FAUVEL, 1895, syn. n.) The roots of this classification are LEE HERMAN'S original tree (HERMAN 1970) and character list. HERMAN did not present a character matrix; his data are inferred from the tree and character table. But even based on these data, the basal branch of the tree is at an incorrect place. It should also be noted here that HERMAN'S phylogeny (HERMAN 1970) was not a quantitative phylogenetic analysis; it was merely a Hennigian argumentation (still revolutionary in the late 60s). A decade later, NEWTON reanalized the phylogenetic relationships of the basal genera (NEWTON 1982). He thought that HERMAN'S "Character 34" should be reversed in polarity: the well developed second abdominal sternite is in fact the derived character state, not the ancestral one. This change had a profound effect: the tree was re-rooted. A related character, the presence of a long articulating membrane connecting the second and third stemites, is correlated with the presence of the well-developed second sternite. All non-oxytelines have the second sternite fused to the third and is narrowed and more lightly sclerotized than that. The taxa he studied (Syntomium, Oxypius, Euphanias) all had the character state possessed by other staphylinid subfamilies, i.e., the reduced, fused second sternite. Another
