Matskási István (szerk.): A Magyar Természettudományi Múzeum évkönyve 98. (Budapest 2006)

Makranczy, Gy.: Systematics and phylogenetic relationships of the genera in the Carpelimus group (Coleoptera: Staphylinidae: Oxytelinae)

of stiff setae. Tarsal segmentation [28] 3-3-3 with no pseudosegment [29] and basal articles [30] compressed or sometimes uncompressed. Ventral setae [31] modified to form tarsal lobes, last tarsomcre [32] only with sparse setae. - Elytra. Elytra [33] without puncture-rows, elytral suture [34] parallel, epipleural ridge [35] present. - Abdomen. Abdomen [36] with two pairs of laterosclerites. Intersegmental membrane [37] without brickwall pattern. Second sternite [38] fully developed, first sternite [39] completely absent. Tergal basolateral ridges [40] absent, carina [41] not present on any sternites. Fimbriate edge [42] on tergite VII unmodified or modified into comb (in which case the fimbriate edge itself is lost). Tergum X [43] sometimes divided by transversal line, its dorsal struts [44] not developed, its basolateral parts [45] not fused to tergum IX. - Secondary sexual characters. Macrocephaly [46] sometimes present, macrothoracy [47] absent. Sternum VII sometimes with spines [48] or with a row of modified setae [49]. The apical edge of sternite VIII [50] shows sexual dimorphism, but bears no tubercles. - Male terminalia. Sternum IX [51] present, with or without setae [52] on apex. - Male genitalia. Aedeagus median lobe [53] bulb-like or laterally flattened, internal sclerites [54] present, symmetrical/paired. Apical opening [55] sometimes well developed (in this case not modified into clasping structure), but often simplified (truncated) or completely membraneous (amorphous). Median face membranous, apically bearing sclerotized edge or contains embedded sclerotized plates [56], without apicomedial hook [57]. Paramercs [58] often wrapping around the middle of median lobe, sometimes with extra lobe [59], sometimes with membranous region [60], with none, or sometimes one or two setae [61] each. Without visible pump and flagellum [62]. - Female terminalia. Female genital appendages [63] lost. Sclerites (possible remnant of sternum X) [64] appear as small ring. - Female genitalia. Spermatheca [65] sclerotized. Receptacle [66] divided, not associated with visible or invisible [67] spermathecal gland, tubular portion [68] sometimes penetrating distal bulb, umbilicus [69] present or absent. Sclerotized "tube" [70] sometimes can be found. Diversity and distribution — Thinodromus is distributed worldwide, contains approximately 200 species and exhibits the most overwhelming diversity within this group of genera. Those species where habitats are known live in periaquatic situations, but always associated with running waters. The specimens often hide under stones and unlike other oxytelines move (and fly away) rather fast, making their collection a challenge. A range of species prefer living in mosses at streams. Trogactus SHARP, 1887 Trogactus SHARP, 1887 (type species: Trogactus championi SHARP, 1887) Description - Medium small to large sized (1.9-6.8 mm), usually dark brown, the smallest are medium to light brown (even if fully coloured). Body with sparse, weakly developed pubescence especially on the abdomen (besides few longer hairs in angles). Antennae extremely long and slender (mid-antennal segments always more than 3 times longer than broad). Elytra parallel-sided, often with tubercles or projections near the postero-lateral corners, temples developed, abdomen usually very narrow at base, broadest at (or around) the 6th segment. The difference in the formation of sternite VIII (not broadly rounded) helps recognizing the males. - Head. Clypeus [1] well developed, epistomal sulcus [2] present. Eyes [3] not extending to the ventral portion of the head. Gular sulci [4] confluent at least anteriorly. Head [5] without midline keel. The mola [6] on the mandibles separated, well developed. Labrum [7] divided, transverse [8]. Setation behind front of labrum [9] in middle