Matskási István (szerk.): A Magyar Természettudományi Múzeum évkönyve 98. (Budapest 2006)

Makranczy, Gy.: Systematics and phylogenetic relationships of the genera in the Carpelimus group (Coleoptera: Staphylinidae: Oxytelinae)

broader tip of terminalia). - Head. Clypeus [1] well developed, epistomal suture [2] present. Eyes [3] not extending to the ventral portion of the head. Gular sulci [4] confluent, but not on anterior portion. Head [5] without dorsal midline keel. The mola [6] on the mandibles separated, well developed. Labrum [7] divided, transverse [8]. Setation behind front of labrum [9] in middle much smaller, irreg­ular or missing. Dorsal setae [10] on epipharynx absent. Hypopharynx [11] undivided, without coriaceous field [12] on ventral side. Lateral row of bulbous setae [13] present. Front of hypopharynx [14] not entirely covered with setae, no setae [15] on mid-line. Median sclerotized plate of hologlossa [16] anteriorly enlarged into a plate with central stick continuing till end. Last palpomere of maxillary palp [17] fully formed, strongly bulbous basally, digitiform sensillae [18] present on last palpomere, arranged [19] in parallel fashion. Labial palp [20] not hairy. Antennae [21] straight and [22] moder­ately elongated. - Thorax. Prosternai process [23] pointed, scutellum [24] without pubescence, hypo­mera [25] not exposing protrochantins. - Legs. Tibia [26] with mid-tibial spur(s) and [27] spines or rows of stiff setae. Tarsal segmentation [28] 3-3-3 with no pseudosegment [29] and basal articles [30] compressed. Ventral setae [31] modified to form tarsal lobes, last tarsomere [32] only with sparse setae. - Elytra. Elytra [33] without puncture-rows, elytral suture [34] parallel, epipleural ridge [35] present. - Abdomen. Abdomen [36] with two pairs of laterosclerites. Intersegmental membrane [37] without brickwall pattern. Second sternite [38] fully developed, first sternitc [39] completely ab­sent. Tergal basolateral ridges [40] absent, carina [41] not present on any sternites. Fimbriate edge [42] on tergite VII unmodified. Tergum X [43] not divided, its dorsal struts [44] not developed, its Table 1. Data matrix of character states for the Carpelimus group and outgroup taxa. Inapplicable states are shown as "-" Platydeleaster bitnacu 1 a tu Deleaster dichrous Coprophilus longicollis Manda inandibularis Planeustomus palpalis Crassodemus foraminosus Aploderus caelatus Aploderus schweigeri Paraploderus parcepunctus Oxytelus laqueatus Anotylus seticornis Bledius talpa Eppelsheimius miricollis Eppelsheimius pirazzolii Sciotrogus opacus Thinobius apfeibecki Thinobius longipennis Neoxus crassicornis Blediotrogus guttiger Pareiobledius alutellus Teropalpus luniger Ochthephilus omalinus Ochthephilus monticola Carpelimus similis Carpelimus exiguus Carpelimus elongatulus Carpelimus bifidus Xerophygus pallipes Xerophygus hreblayi Mimopaederus insularis Thinodromus priorus Thinodromus dilatatus Thinodromus corvinus Thinodromus caloderinus Thinodromus skottsbergii Thinodromus juanfernandez Thinodromus luteipes Thinodromus diffusus Thinodromus chagosanus Thinodromus thoracicus Thinodromus indicus Thinodromus schi 1lhammeri Thinodromus singularis Trogactus ocellatus Trogactus godmam 123456789 000000000 000100000 010000000 101001001 101002001 000100102 000100102 000100102 000100102 010100002 000100002 000100102 000100003 000100003 000100012 000100012 000100012 000100012 000200102 000100102 000000102 000100002 000100002 000100102 000100102 000100102 000100102 000100102 000110102 000100102 000100102 000100102 000100102 000100102 000100102 000100102 000100102 000100102 000100102 000100102 000100102 000100102 000100102 000100102 000100102 0123456789 0000000000 0000000000 0000100001 0000000000 0000000001 1101110001 1001000002 1001000002 1101000002 1001000302 1101000302 0010110002 0010010001 0010010002 111000133­1010001202 1010001202 1010001202 1011001102 1011000102 1011001102 1000002102 1000001102 1010002202 1010002202 1010002202 1010101202 111111233­101111233­1010101102 10L0102202 1111101202 1010001322 1110101322 1110101202 1110101202 1110001202 1111002312 1110102202 101010233­1010102322 1010102322 1010102322 111010133­1110101312 0123456789 0000000000 0000000000 0000000000 0000011000 0000011011 1000011010 0000001020 0000011020 0000011020 0000011020 0000010020 0100010010 01OOOOO0O0 0100000000 0000001130 0000101130 0001101130 0000111130 0OOOO11O0O 0000011010 0000011100 0000001100 0000001100 0000011120 0000010120 0000011120 0000011120 0000011000 0000011000 2000011100 0000011100 0000011100 2000011100 2000011100 0000010100 2000011100 2000011100 0000011100 0000011100 2000011100 2000011100 2000011100 2000011100 2010011100 2010011100 0123456789 0000000000 0001000000 0000001000 0001001011 0000001111 0101001111 0101001111 0101001111 0101001111 0101001111 0101001111 0001101111 0001101111 0001101111 -101011111 -101111111 -101111111 -101101111 1101001111 1101001111 0101001111 1101001110 1101001110 1101001111 1101001111 1101001111 1101001111 1101001111 1101001111 1101001111 1101001111 1101001111 1101001111 1101001111 0101001111 0101001111 1101001111 1101001111 1101001111 1101001111 1101001111 1101001111 1101001111 1111001111 1111001111 0123456789 0000000000 0000000000 0000100100 0000000000 0000000000 0000201000 0000201000 0020201000 1000200000 1000201000 2000211000 0000200000 0000201000 0000201000 1000200000 1000200000 1000200000 1000200000 0000200000 0000201000 0010200000 0000200000 0010200000 0000200000 0000201000 0000200000 0000200000 0010211000 0000211000 0100200000 0000200000 0000200000 0000200000 0001200000 0000200000 0020201000 0000200000 0000200010 0000200000 0000200001 0000200000 0000200000 0000200000 0100200000 0100200000 0123456789 0000000000 0000100000 0000240000 0000140000 0000040000 0000200000 0000200000 0000200000 0010231100 11-0210100 01-0200000 0000210000 0000040000 0000040000 0010211000 0000200000 0000240000 0000240000 0000210000 0000210000 0000230000 0000100000 0000000001 01-0210000 01-0210000 01-0210000 01-0210001 0000200000 0000210000 0000230000 0011210000 0010222010 0010202011 0001220000 0011220000 0001220000 0001220000 0001200000 0000210000 0000211010 0000220011 0000210001 0000210000 0000210000 0000210000 0123456789 0 000000-0-- 0 000020-0-- 0 000010-0-- 0 0000211000 1 0000210000 0 0001011000 1 0104010100 0 0104010100 0 0304211100 0 0004010100 0 0102011101 0 1201211000 0 0000210000 0 0000210000 0 0000111000 0 0002111001 0 0013210001 0 0013211001 0 0001211100 1 0001211000 1 0000211100 0 000010-0-- 0 100010-0-- 0 1104111100 0 0104111100 1 1104111100 0 0104211000 0 0111211101 0 0011211101 0 0000111100 0 0204211100 1 1004111100 0 1004011000 0 1004111010 0 1004211101 0 1104211101 0 1104211101 0 0104011110 0 0104111110 1 1204111100 0 1004211100 0 1104211000 0 1004111000 0 1004210000 0 1004210000 0

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