Matskási István (szerk.): A Magyar Természettudományi Múzeum évkönyve 97. (Budapest 2005)

Bálint, Zs.: A review of the Neotropical hairstreak genus Annamaria with notes on further genera (Lepidoptera: Lycaenidae)

The male of A. Columbia: All the genera closely related to Evenus (see Discussion below) are dimorphic sexually, but ventral markings of the sexes are qualitatively identical. Hind wings sometimes are dissimilarly tailed, for example Aveexcrenota anna (DRUCE, 1907) JOHNSON et CONSTANTINO, 1997 and Paraspiculatus catrea (HEWITSON, 1874), BÁLINT et MOSER, 2001. The male fore wing ventra could have structural colouration as in Atlides polybe (LINNAEUS, 1767) HÜBNER, 1819 or Denivia hemon (CRAMER, 1775) JOHNSON, 1992. Male wing dorsa are more bright, females are darker blue with wide distal margin or even brown {Macusia lineage). Their ventral wing is qualitatively different in hue or quantitatively distinct in colour (D. hemon, Margaritheclus danaus (FELDER et FELDER, 1865) BÁLINT, 2002, Theritas drucei LATHY, 1930, but their patterns are always identical qualitatively. Consequently I presume based on analogies of congeneric taxa and other large eumaeines that the male wing ventrum of A. Columbia similarly patterned as the the holotype female. A. Columbia male hind wing should have three hind wing tails, the fore wing distal margin should be somewhat shorter resulting slightly narrower shape. The dorsal colouration is certainly somewhat lighter bluish green with fore wing discal scent pad and subdiscal patches and narrow black marginal border. The ventral wing ground colour should be somewhat lighter, fore wing probably possessing structural blue medial area. There is no evidence for that the phenotypes of A. rhaptissima and A. Columbia would represent the same biological species. Figs 23-24. Brevianata saphonota (CONSTANTINO, SALAZAR et JOHNSON, 1993), comb, n., female, Ecuador, Esmeraldas (coll. P. BOYER): 23 = dorsal view, 24 = ventral view. (Images taken with the same magnification as Figs 4-5)