Matskási István (szerk.): A Magyar Természettudományi Múzeum évkönyve 92. (Budapest 2000)

Ronkay, L.: Revision of the genus Lophoterges Hampson, 1906 (s. l.) (Lepidoptera: Noctuidae, Cuculliinae). Part I

strongly from the true Lophoterges species in several external and genital features. The detailed study of these taxa revealed the facts that 1 ) these two species belong to two distinct genera, which are not only rather remote from the Lophoterges-Yme but even the correct tribal (subfamilial sensu POOLE 1995) placement of whose is problematic; 2) 'L. taiwana ' represents a group of closely related taxa, two of them are yet undescribed. These two genera, Atakterges gen. n. and Speidelia gen. n., share numerous typical features of Oncocnemidini and Psaphidini, displaying also characteristic autapomoiphies. The features being common with both tribes are the 1 ) long, tubular vesica, covered densely with cornuti, 2) collar producing into acute hood (this character is more typical to some groups of Oncocnemidini but it is present also in some genera which are associated with Psaphidini, e.g. Pleromella DYAR, 1921; Provia BARNES et MCDUNNOUGH

, 1910). The characters shared with Psaphidini are 1) irregular spining of fore tarsi, although spines not very strong; 2) early spring (and late autumnal) flight period of adults. The characters shared with Oncocnemidini are 1) female genitalia with strongly bilobate bursa copulatrix; 2) ductus seminalis arising from apical end of cervix (appendix) bursae; 3) male genitalia with more specialized clasping apparatus; 4) presence of large frontal prominence in one genus; 5) male antenna filiform, rather thick, only slightly ciliate; 6) proboscis well developed. The knowledge of larval structures may help to resolve the relationships of the two genera, the early stages of the species, however, are unfortunately unknown. Thus, only the information based generally on the adult morphology can be summarized. The genera Atakterges and Speidelia are considered as primitive Oncocnemidini species, representing ancient stages in the Oncocnemis-Sympistis evolutionary line. The diagnoses and descriptions of the new genera and species are given in the systematic part of the paper. Abbreviations - AKM - Alexander Koenig Museum, Bonn; BMNH - The Natural History Museum (formerly British Museum, Natural History), London; HH -HERMANN HACKER; HNHM Hungarian Natural History Museum, Budapest; NMNS - National Museum of Natural Sciences, Taichung, Taiwan; RL -RONKAY LÁSZLÓ; SS -SHIGERO SUGl; TFRI - Taiwan Forestry Research Institute, Taipei; ZMFK - Zoologisches Forschungsinstitut und Museum Alexander Koenig, Bonn (= AKM).