Matskási István (szerk.): A Magyar Természettudományi Múzeum évkönyve 89. (Budapest 1997)

Hír, J.: Cricetus runtonensis solymarensis ssp. n. (Mammalia, Rodentia) from the Late Middle Pleistocene fauna of Solymár

Cricetus runtonensis solymarensis ssp. n. Holotype. Viscerocranium No. 2. Paratype. Mandibula No. 1. Type locality. Kiskörút Hall in the Ördög-lyuk Cave at Solymár, Buda Mountains, NW from Budapest. Type level. Red coloured argillaceous cave sediment. Age. Lower part of the Toringium after the system of FFJFAR & HEINRICH (1981, 1986, 1990). In the traditional Hungarian vertebrate paleontological stratigraphy (JÁNOSSY 1986) the Solymár phase is the oldest part of the Late Middle Pleistocene. Up to the present the exact correlation of this period with the standard Pleistocene chronostratigraphy (ZAGWIJN 1985) is impossible, but we presume that the Solymár phase refers to the Holsteinian or an other mild climatic period within the Saalien. Etymology. After the name of the settlement, which is close to the locality. Synonyms. Cricetus cricetus inter glaciális KRETZOI, 1946; Cricetus cricetus ssp. JÁNOSSY, 1986. Diagnosis. Large sized hamster subspecies, different from the Lower Pleistocene Cricetus runtonensis runtonensis population of Somssich-hegy 2 in the following statistic morphological features, — the higher frequency of the conelet entostyle in the upper M

1, M2 molars, and of the conelet ectostylid in the lower m2 , m3 molars; — the frequent duplication of the mesolophid in the m2, m3 molars. Among the studied dimensions the LM2, LM3, WaM3, Lml-m3, Waml, Lm2 are significantly larger, the LM1, WpMl, WpM2, LM1/LM2 are significantly smaller in C. runtonensis solymarensis than in C. r. runtonensis. Description. The measurements of the molars are definitely larger, than the corresponding dimensions of the Recent C. cricetus from Hungary (HÍR 1997), Poland (PRADEL 1985) and Germany (FAHLBUSCH 1976), but similar to the metrical data of the C. runtonensis material from Poland (PRADEL 1988) and of the C. runtonensis population of Somssich-hegy 2 in Hungary (Figs 2-7, Tables 1-8). During the detailed comparison of 22 metrical characters of the C. runtonensis materials from Solymár and from Somssichhegy 2 localities 6 characters are significantly larger in the Solymár material (LM2, LM3, WaM3, Lml-m3, Waml, Lm2), 4 characters are significantly larger in the SS2 material . (LM 1, WpM 1, WpM2, LM1/LM2). No significant differnce in the other 12 characters. In the morphological description we will not reinterprete the general odontomorphological picture of the hamsters. We want to give the special features of our material. Ml. There are no remarkable differences in the frequency of the morphotypes among the investigated C. cricetus, C. praeglacialis materials. AOBYOBO is the dominant morphotype everywhere, but in the C. runtonensis populations of Solymár and Somssich-hegy 2, the type AOBYOB 1 is subdominant because the ramification of the posterolophule is more frequent(Fig. 8, Table 9). M2. In the Recent hamster from Hungary and in C. praeglacialis the dominant morphotype is the AOO, but in the C. runtonensis materials (SS 2 and Solymár) AOl is dominant and AOO is only subdominant. In the latter samples the ramification of the terminal part of the posterolophule is much more frequent (Figs 9-10, Table 10).