Matskási István (szerk.): A Magyar Természettudományi Múzeum évkönyve 89. (Budapest 1997)

Bálint, Zs. ; Johnson, K.: New species of Cyanophrys sensu lato from the Andean region of Argentina (Lepidoptera: Lycaenidae)

Remarks. Review of the type data indicates the habitat is mesic upland subtropical forest (in both the northern and southern remnants) except that the Abra Grande locality is today disturbed by agriculture and nothing is known of the original San Pedro de Colaleo habitat. The 1991 Cumbres de San Javier collections were in the vicinity of Villa Nougués (a favorite Hayward locality) and characterized by hilltopping above mesic forest at a broken margin with a roadway. Numerous tropical forest butterfly species occur in this limited region of Tucumán Province; these include, in the Lycaenidae, forest elfins like the Argentine endemics Rhamma amethystina (HAYWARD, 1949) and Pontirama brunea JOHNSON, 1992 and seldom collected forest hairstreaks like Terra carta (HAYWARD, 1949) and Strymon oreala (HEWITSON, 1868) (southernmost record for South America). It is important that lepidopterists not confuse C. octonarius with C. portoena (CLENCH), 1944 (type MCZ, type locality: Cussiluni, Bolivia) a member of the "acaste" species complex (or "superspecies") which has a limited VHW dashed lineal band. Taxa pairs like acaste PRITTWITZ, 1865 / lycimna HEWITSON, 1868 and abnormis CLENCH, 1946 / catharinensis CLENCH, 1944 also belong to this complex and share a distinctive morphology. Geographic distributions of these taxa reflect vicariances related to the Parana River basin, C. portaena being restricted northward in the Bolivian yungas and immediately adjacent areas of Argentina. The complex needs further evaluation as to taxonomic rank. Cyanophrys portaena (or, alternatively, C. acaste portaena, depending on usage) does not have the indented, "figure-8", shape to its white ventral markings and also differs from C. octonarius in the genitalia of both sexes. Etymology. Latin meaning "eight", referring to the distinctive "figure 8"-like markings formed by the postbasal and medial bands of the VHW. For historical consistency we note this species was originally given another manuscript name not used here (JOHNSON 1980: 442, third name in directory of treatments at top of page). Plesiocyanophrys JOHNSON, ElSELE et MACPHERSON, 1993 Plesiocyanophrys was employed by JOHNSON el al. (1993) to denote a clade within Cyanophrys sensu lato characterized by extremely simple genitalic structures. Such uncomplex elements, and those without additional components, were viewed as generally primitive in ELIOT'S (1973) phylogenetic scheme for the Lycaenidae. Members of Plesiocyanophrys usually show only minor differences in wing pattern but genitalic structures can differ markedly. Plesiocyanophrys runa, sp. n. (Figs 7, 18) Diagnosis. This species is currently known only from the limited remnant subtropical forests of Catamarca Province, Argentina. All southern South American Plesio-