Matskási István (szerk.): A Magyar Természettudományi Múzeum évkönyve 87. (Budapest 1995)

Dulai, A.: Middle Miocene (Badenian) lunulitiform Bryozoa from Szob (Börzsöny Mts., Hungary)

SYSTEMATIC DESCRIPTIONS Class: Bryozoa EHRENBERG, 1831 Subclass: Gymnolaemata ALLMAN, 1856 Order: Cheilostomata BUSK, 1852 Family: Cupuladriidae LAGAAD, 1952 Genus: Cupuladria CANU et BASSLER, 1919 Cupuladria vindobonensis BALUK et RADWANSKI, 1984 (Figs 4-10, 23, 27-28) Cupularia canadensis BUSK: MANZONI 1877: 72, PI. 17, Figs 56a-c. Cupuladria canariensis (BUSK): LAGAAIJ 1952: 33, PI. 2, Figs la-b; LAGAAIJ 1953: 15, Pl. 1, Figs la-b; BUGE 1957: 139, Pl. 9, Fig. 5, Pl. 10, Fig. 3; LAGAAIJ 1963: 172, Pl. 25, Figs la-b, 3-5; ANNOSCIA 1963: 225, Pl. 9, Fig. 1, Pl. 10, Fig. 1, Pl. 11, Figs la-b, Pl. 12, Figs la-b; CHEETHAM & SANDBERG 1964: 1021, Figs 11, 13; COOK 1965b: 197, Figs la-f, Pl. 3, Fig. 4; BUGE 1973: 36, Pl. 6, Figs 1-2; CADÉE 1975: 323, Pl. 3d; BALUK & RADWANSKI 1977b: 143, Pl. 1, Figs 1-6, Pl. 2, Figs 1-5, Pl. 3, Figs. 1-2; CADÉE 1977: 45, Pl. 1, Fig. 3 Cupuladria canariensis canariensis (BUSK): CADÉE 1979: 446, Figs lc-d. Cupuladria canariensis cavernosa CADÉE 1979: 445, Figs le-g, Fig. 4; BISHOP & HAYWARD 1989: 8, Figs 18-21. Cupuladria vindobonensis BALUK & RADWANSKI 1984a: 22, Pis 2-3, PI. 9, Figs 1-2; BALUK & RADWANSKI 1984b: 243, Pis 1-4; REGUANT 1993: p.127. Material: Numerous fragmentary specimens. Dimensions: length of zooecia 0.637±0.0324 mm (range: 0.59-0.70 mm), width of zooecia 0.276+0.013 mm (range: 0.22-0.31 mm), length of vibracular opesia 0.206+0.01 mm (range: 0.18-0.27 mm), number of pores per basal sectors 8.56± 2.99 (range: 2-17) (see details in Tables 1 and 3). Description: The discoidal, flattened cone-shaped zoaria are free in adult stages. Zooecia form a single layer, in which vibracula alternate with zooecia in the same radial series. The asymmetrical vibracula aie distal to each zooecia. The shape of zooecia is rounded rectangular. Central zooecia arc open, without closures by a calcareous lamina. Vestibular arch is not developed. The cryptocyst is simple, without any denticles. Ovicells are absent. Opesia are deprimed laterally. The number of basal kenozoidal chambers are between 1-5. The kenozoecial layer is about as thick as the zooecial layer. The basal surface of the colony is sectored by radial and tangential grooves. The majority of the basal sectors are long, with pores ranging usually between 5-13 and their maximum range observed 2 to 17 in a sector. Remarks: The family Cupuladriidae was described by LAGAAIJ (1952) to include only the type genus Cupuladria. Later Discoporella also was classified to family Cupuladriidae by CHEETHAM & SANDBERG (1964), COOK (1965a) and BUGE (1973) but recently BALUK & RADWANSKI (1984a) returned to the original interpretation and Discoporella was placed into a new family (Discoporellidae BALUK et RADWANSKI). According to COOK (1965a) three species belong to her Cupuladria canariensis group, i.e. C. canariensis (BUSK), C. monotreinata (BUSK) and C. biporosa CANU et BASSLER). This group was supplemented by CADÉE (1975) and BALUK & RADWANSKI (1984a, b) with two later described species (C. surinamensis CADÉE, C. vindobonensis BALUK et RADWANSKI). A new subspecies was described by CADÉE (1979) for European Miocene-Pliocene Cupuladria canariensis (C. canariensis cavernosa CADÉE) on the basis of the number of kenozoecial chambers in the basal pad. Recent C. canariensis (BUSK) has only one kenozoecial chamber in a row but in the fossil forms there are more than one chambers in a row. In CADÉE'S interpretation C. canariensis canariensis (BUSK) has 5.6-14.2 (1-28) and C. canadensis cavernosa (CADÉE) has 4.5-8.1 (1-28) pores at basal sectors. However, according to BALUK & RADWANSKI (1984a, b) the number of chambers depends on the thickness of the basal pad and is ascribable to ecological factors without any taxonomic value. Instead of this a new species was described (C. vindobonensis BALUK et RADWANSKI)