Matskási István (szerk.): A Magyar Természettudományi Múzeum évkönyve 85. (Budapest 1993)
Nesemann, H.: Identification key to the Hungarian leeches of the subfamily Trochetinae Pawlowski, 1954, with notes on systematics of the subfamily Erpobdellinae Blanchard, 1894 (Hirudinea)
Diagnosis: Large leeches up to 100 mm length, annulation formula of somites: cl, c2, b2, a2, b5 (often subdivided in c9, clff), ell, d23, d24 (Fig. 49). Long and rostelloid oral sucker with small pseudognaths (Fig. 46). Atrium restricted on somite XII, reaching from XII b2/a2 to XII cl/c2. Atrium very small with not coiled cornua (Figs 47-48). Male gonopore in the furrow XII b2/a2, female in XII b5lcll. Colour variable, in general black or dark reddish brown, always with one pair of dirty paramediane stripes. Light greyish and small spots tranversal on each annulus, irregularly arranged in two rows (Fig. 49). The body of this species resembles T. pseudodina, which is quite common in western Europe. It differs from T. pseudodina and T. cylindrica in the small size and form of the atrium, much smaller pseudognaths and dorsal colouration, which is similar to Dina punctata and Dina stschegolewi. Distribution: Tributaries of the upper Rába river basin in Hungary and Austria (southern Burgenland), most probably in adjacent waters in Slovenia (SKET, pers. comm.). All of them were western tributaries of the Pliocene Slavonian lake, a relict water of the former Pannonian sea. Habitat: Semiaquatic on the banks of small rivers and streams. IDENTIFICATION KEY TO THE SPECIES 1 (2) Smaller to medium-sized leeches with cylindrical or slightly flattened body, five annuli per somite, all of them with the same width. Small pseudognaths, suctoroid oral sucker. - Erpobdella DE BLAINVILLE, 1818 (see PAWLOWSKI 1948, Soós 1963b, 1967) 2 (1) Medium-sized to large leeches with cylindrical preclitellar region and more flattened posterior part of the body, often with lateral keels. Annulus b6 always widened. Five to nine annuli per somite. Ejaculatory ducts of male genital system with pre-atrial loops or atrial cornua in somite XI. - Trochetinae PAWLOWSKI, 1954 3 (7) Annulus b6 not subdivided or homonomously subdivided in rings of the same width (e.g. ell, cl2). - Dina R. BLANCHARD, 1894 4 (5) Annulus b6 sometimes with a shallow furrow subdivided in ell, cl2. Reddish brown colour with dark paramedian stripes. Gonopores separated by 2 or 2 1/2 annuli, male gonopore in the furrow of XII b2/a2, female in XII b5fb6 or on XII b6. Suctoroid oral sucker, segments I-II short Dina lineata (O. F. MÜLLER, 1774) 5 (4) Dark blackish ground colour, small yellowish spots tranversal on each annulus, irregularly arranged in two rows, one pair of dark paramedian stripes. Gonopores separated by 2 or 1 1/2 annuli. Rostelloid oral sucker, segments I-II widened Dina punctata JOHANSSON, 1927 6 (5) Dark blackish ground colour, large yellowish spots transversal on each annulus, irregularily arranged in one row, on b6 in two rows. Lacking dark paramedian stripes. Gonopores separated by 2 annuli. Rostelloid oral sucker, segments I-II widened Dina apathyi GEDROYC, 1916 7 (3) Annulus b6 widened and heteronomously subdivided in one tertiary (ell) and two quartary rings (d23, d24). Five short and three wide or seven short and two wide annuli per somite. Prominent pseudognaths, rostelliod oral sucker with strongly developed rostellum, widened segments I and II. - Trocheta DUTROCHET, 1817 8 (11) Large unicoloured leeches, neither dark stripes nor yellowish spots. Very large pseudognaths.
