Matskási István (szerk.): A Magyar Természettudományi Múzeum évkönyve 85. (Budapest 1993)

Papp, L.: Three new milichiid species (Diptera, Milichiidae) from Hungary

Head (Fig. 9) higher than long. Ocellar triangle extended to a shining interfrontal triangle reaching nearly to lunule. Lunule very short, facial keel low and not sharp, mouth edge slightly protruding in profile (Fig. 9). Gena half as broad as flagellomere. Both ors pairs exclinate, both ori pairs inclinate; a short (0.052 mm) additional ors between posterior ors and vti; vti extremely long, poc pair long, strong and inclinate. Flagellomere comparatively longer than in Xenophyllomyza deserticola (Fig. 8, cf. Fig. 1 of OZEROV 1992): length/width 0.147mm/0.112mm, with long cilia. Arista short, 0.259 mm, its cilia as long as on flagellomere. Whole length of proboscis 0.86 mm. Apex of palpi with 5 straight bristles. Thoracic chaetotaxy: 1 h, 2 np, 1 prsut, 1 sa, 2 pa, 2 dc, 1 prsc, 2 sc. Anterior dorsocentral caudal to supraalar and less than half as long as posterior one. Apical scutellars widely divergent. Proepisternum with a minute bristle, anepisternum and anepimeron bare. Katepistemum with 1 very strong bristle. Legs brown, short but robust, tarsi light brown. Tarsomeres slightly dorso-ventrally flattened. Mid tibia with a strong ventroapical, otherwise legs without characteristical bristles. Membrane of wings with some light brownish hue, costal and radial veins greyish ochreous, other veins indistinct yellowish. No hind crossvein (dm-cu) nor even a vestige of a vein there (Fig. 10). Veins R2+3 and R4+5 nearly parallel and close to each other. t» crossvein proximal to Rl break of costa (Fig. 10). Anal vein distinct on a section of c. 0.2 mm as a faint line. Halteres black. Abdominal terga with dark grey microtomentum. Male genitalia not studied. Paramyia hungarica sp. n. is the first known species of the genus Paramyia WILLISTON, 1897 from the Palaearctic. Xenophyllomyza OZEROV, 1992 (type-species: X. deserticola) described most recently from Turkmenistan seems related (or even congeneric), but this question must be cleared in a subsequent revision of the species of Paramyia. In the collection of the HNHM there are specimens of 2 undescribed species from the Neotropical region and another species (inconspicua MALLOCH or a new species) from Viet Nam (inconspicua hitherto known from Java only). SABROSKY (1989) listed "Unidentified spp." from Australia. That revision seems even more interesting since it has been found that Paramyia and Aldrichiomyza HENDEL are related. Their shared synapomorphies are: costa extending to R4+5, proboscis very long, no vibrissae, veins R2+3 and R4+5 parallel and close to each other, veins R4+5 and M divergent. Aldrichiomyza and Xenophyllomyza share also the features that anterior ors are proclinate and both possess 2 pairs of katepisternals. The differentiating characters of Paramyia are its normal pubescent arista, and the loss of dm-cu crossvein (and I oxide pairs). The only synapomorphies for the species of Aldrichiomyza are: their thick arista with dense thick short setae, more dorsocentral pairs (incl. a presutural pair) and the retention of dm-cu crossvein. It seems indispensable to study the details of male genitalia in order to clear up relations among those three genera. References HENDEL, F. (1913): H. Sauter's Formosa-Ausbeute. Acalyptrate Musciden (Dipt.) - Supplia Ent 2: 77-112. HENNIG, W. (1937): 60a. Milichiidae et Carnidae. - In: LINDNER, E. (ed.): Die Fliegen der paläarktischen Region, VI, 1. Schweitzerbart'sehe, Stuttgart, 91 pp. OZEROV, A. (1992): Xenophyllomyza deserticola gen. et sp. n. (Diptera, Milichiidae) iz Turkmenistana. (Xenophyllomyza deserticola gen. et sp. n. (Diptera, Milichiidae) from Turkmenistan.) - Vestn. zool 1992 (2): 18-20. PAPP, L. (1978): 72. család: Milichiidae-Pákosztoslegyek. - In: DELYNÉ DRASKOVTTS, Á. & PAPP, L.: Taplólegyek-Gabonalegyek - Odiniidae-Chloropidae. Magyarország Állatvilága (Fauna Hungáriáé) 15, 9. Akadémiai Kiadó, Budapest, pp. 9-32. SABROSKY, C. W. (1965): Family Milichiidae (Phyllomyzidae, Carnidae). - In: STONE, A. et al. (eds): A Catalog of the Diptera of America north of Mexico. Washington, pp. 728-733. SABROSKY, C. W. (1973): 75 Family Milichiidae (Phyllomyzidae, including Carnidae). - In: PAPAVERO, N. (ed.y.A Catalogue of me Diptera of me Americas south of the United States. Sao Paulo, pp. 75.1-75.12.