Matskási István (szerk.): A Magyar Természettudományi Múzeum évkönyve 83. (Budapest 1991)

Kvaček, Z. ; Hably, L.: Notes on the Egerian stratotype flora at Eger (Wind Brickyard), Hungary, Upper Oligocene

FAGACEAE Quercus rhenana (KRÄUSEL et WEYLAND, 1950) KNOBLOCH et KVACEK, 1976 (Plate VI: 2,3) Fragmentary lanceolate leaves c. 3 cm wide and more than 10 cm long, entire-margined, acuminate at the apex, with stout midrib passing into a thick petiole, and regulary looping dense secondaries (at 60°-80° angles to the midrib) were identified at BP d&Apocynophyllum sp. (BP 70.102.1, 70.451.1), or Lithocarpus sp. II sensu ANDREÁNSZKY (1966, textf. 35, BP 67.72.1). We do not hesitate to identify them with Q. rhenana, a common member of Late Oligocene-Lower Miocene swamp forests. All specimens come from EWU. cf. Lithocarpus saxonicus WALTHER et KVACEK in KVACEK et WALTHER, 1987 (Plate VII: 4) Rare lanceolate entire-margined leaves about 4-5 cm wide and up to 20 cm long are characteristic by camptodromous fagaceous venation, secodaries very regular, steep (at an angle of 30° at the leaf apex) and distinct percurrent tertiaries directed perpendicularly to the midrib or nearly so. They partly correspond to the entity called by ANDREÁNSZKY (1966)L/£seű cf. term (L.) MERR. (1. c. textf. 12, MM 7394, 64.176.1, non f. 13 - see ? Fagaceae gen.) and are confined to EWM. In their leaf form they well match with a newly described Lithocarpus saxonicus from the Oligocène of Germany (KVACEK & WALTHER 1987) but differ by the less oblique course of tertiary venation. Dryophyllum callicomifolium (ANDREÁNSZKY, 1962) comb. n. (Plate VI: 4) B a s i o n y m : Castanopsis callicomaefolia Andreánszky, 1962, 221. pl. 2. f. 2-4. pl. 3. f. 1 Synonyms: Quercus agriensis ANDREÁNSZKY, 1962,224. pl. 3. f. 2 Quercus crassipetiolata ANDREÁNSZKY et KOVÁTS in ANDREÁNSZKY 1966.64. textf. 50-53 Quercus legányi ANDREÁNSZKY et KOVÁTS in ANDREÁNSZKY 1966.52. textf. 36,37. H o I o t y p e : ANDREÁNSZKY 1962, pl. 2, f. 4 (missing at MM) - EWL. - Paratypes: MM 61.793.1, ANDREÁNSZKY 1962, pl. 2, f. 3 - EWL, MM 61.817.1, ANDREÁNSZKY 1962, pl. 2. f. 2 - EWL. This leaf form peculiar by its slender form and regular, blunt and partly S-shaped teeth is a common element in EWL, much rarer in EWM. We have not found any essential difference between the three ANDREÁNSZKY's species: Castanopsis callicomifolia, Quercus crassipetiolata and Q. leganyi and unite them under one entity. Since there is no real basis for a generic determination we suggest to accomodate it into the form genus Dryophyllum in a common sense (see KVACEK & WALTHER 1989a, b). It is not excluded that this Late Oligocène species is a descendent of Eotrigonobalanus furcinervis (ROSSM.) WALTHER et KVAÖEK but it decidedly differs in its much slender and less variable leaf form. Not knowing the epidermal characters we can hardly decide this question. Callicoma egedensis ANDREÁNSZKY et NOVAK (1957, pl. 2. f. 3) from Kiseged does not differ in its leaf form and dentation (see also ANDREÁNSZKY 1966: 88) but the type (BP 83.256.1-5181) shows no details of venation except secondaries, the surface is covered by false contours left by fossilisation processes and thus there is no serious basis to employ this basionym for D. callicomifolium, Quercus agriensis in our opinion represents only an aberrant small leaf of this species. We found also almost entire-margined forms (e.g. MM 64.129.1) similar to that figured by ANDREÁNSZKY (1962, pl. 3. f. 1). ? Trigonobalanopsis rhamnoides (ROSSMÄSSLER, 1840) KVACEK et WALTHER, 1988 (Plate VI: 5) A single specimen described as Quercus tenuipetiolata ANDREÁNSZKY (1962, 222. textf. 2, BP 83.278.2) and its counterimprint (BP 26030) match well with this well-known subtropical element of European Tertiary, mainly of eucamptodromous venation, better seen in the counterimprint and oblique course of tertiary