Matskási István (szerk.): A Magyar Természettudományi Múzeum évkönyve 80. (Budapest 1988)

Roháček, J. ; Papp, L.: A review of the genus Paralimosina L. Papp (Diptera, Sphaeroceridae), with descriptions of ten new species

pattern similar to the Af-shaped mark of Paralimosina species, forms a sister-group to Paralimosina. However, Biroina species, which also occur in the Oriental Region and not only in the Australian Region as stated by RICHARDS (1973), have a simple hypandrium and different aedeagal morphology and t

2 chaetotaxy. Another taxon related to Paralimosina seems to be Limosina monorbiseta (DEEMING, 1969) from Nepal (L. PAPP, unpublished). More precise data on the postabdominal structures of both the above taxa are needed to evaluate their affinities to Paralimosina. Both Hackmaniella L. PAPP, 1979 and Nipponsina L. Papp, 1982 were based on apterous species having some striking modifications reflecting their terricolous life-habits. Analysis of their genitalia confirms that they belong to Paralimosina and therefore Hackmaniella and Nipponsina are synonymized here. With the 10 new species described below, the genus Paralimosina now comprises 20 species in the Palaearctic and Oriental Regions. No Paralimosina species is known from the New World. Leptocera (Scotophilella) levigena SPULER, 1925, assigned to Paralimosina by ROHÁCEK (1983) on the basis of its original description, belongs to the genus Kimosina ROHÁCEK, 1983 (S. A. MARSHALL, pers. comm., 1986). On the other hand, Paralimosina is represented in the Afrotropical Region by several unnamed species (examined) and most probably also in the Australian Region. Judging from the species diversity of Paralimosina discovered in SE Asia, further undescribed species can be expected there and in adjacent areas. The gaps in our knowledge of some of the regional faunas precluded construction of a reliable phylogenetic hypothesis for Paralimosina. However, the relationships (if recognized) are discussed under each species reviewed here (in alphabetical order). KEY TO THE IDENTIFICATION OF PARALIMOSINA SPECIES 1 Apterous species without haltères and with broad abdomen 2 Wings (sometimes shortened) and haltères present; abdomen not strikingly broad 3 2 (1) M-shaped mark on frons well developed, velvety, dark brown. Abdomen with shining, hardly tomentose, terga with long pale setae. Male genitalia (Figs 34-38), female postabdomen (Figs 39-42, 45, 46) P. ceylanica (L. PAPP) M-shaped mark not developed. Body, including abdomen, with dense greyish brown microtomentum, dull. Abdominal terga with short, dark setae. Male genitalia (Figs 124-127), female postabdomen (Figs 128-132) P. sexsetosa (L. PAPP) 3 (1) M-shaped mark not developed; frons unicolorous. Fore tarsomeres 2-5 white to yellowish white, contrasting with dark mt

1 4 — M-shaped mark distinct, usually velvety, dull (rarely subshining — P. indica sp. n.), black to brownish orange; frons variegated. Fore tarsus unicolorous 6 4 (3) Frons fuscous ochreous. Wings somewhat shortened, with diffuse dark brown spots. Large species (body length 2.8 mm), with thickened male /^and /

3

. Gonostylus (Fig. 82) with only thin long setae on posterior lobe. Male genitalia (Figs 61-64). Female unknown P. gigantea sp. n. Frons dark brown to black. Wings shortened or normal, without darker spots. Smaller species (body length less than 2.4 mm), male with slightly thickened f

y and /

3

. Gonostylus with a robust seta on posterior lobe 5 5 (4) Only 1 ors and 2 ifr (anterior minute). /

3 and r

3 unicolorous, pale ochreous brown. Wings fully developed though sometimes narrowed and slightly shortened. Male genitalia (Figs 65-68), female postabdomen (Figs 70, 71, 74, 75) P. icaros sp. n.