Kovács I. (szerk.): A Magyar Természettudományi Múzeum évkönyve 78. (Budapest 1986)

Ronkay, L.: On the taxonomy and zoogeography of some Palearctic and Indo-Australian Plusiinae (Lepidoptera, Noctuidae)

darker with some ochreous scales. Cilia whitish with dark spots. Alar expanse 33-36 mm, length of fore wing 16-18 mm. Male genitalia (Figs 1-2): Uncus long and slender, tegumen wide and moderately high, fultura inferior long and slightly pointed. Vinculum very long and less sclerotized. Valvae arcuate, cucullus narrow, outer margin dentated, corona absent. Costal margin strongly curved, clavus very long and thin, sacculus with two basal appendages and a large distal lobe; these lobes asymmetric, right lobe more elongate and larger. Aedoeagus moderately long, slightly curved, ventral sclerotized ribbon strong, basally furcated. Vesica everted ventrally, basal part with a spinulose lamina and a slender, arcuate cornutus on ventral side. Apical part of vesica with two diverticles, one of them with fine spicules on upper side. Ventral surface with two rows of strong cornuti, caudal surface with a globular diverticle covered with fine spinules. Female genitalia (Fig. 25) : Papillae anales less chitinized, gonapophyses long and thin. Ostium bursae granulate, ductus bursae wide. Apex bursae with sclerotized crests, corpus bursae narrow at middle, fundus wider, rounded; signum large, round. Distribution : Korea, China, Japan (Sugi, pers. comm.), Amur Region (USSR). Specific differences and systematic position — The new species with its morphological characters shows an interesting intermediate status between the related species, M. confusa (STEPHENS, 1850) and M. crassisigna (WARREN, 1913). Externally M. hybrida is very similar to M. crassisigna (Plate: 3) with its large, bilobate or conjoined stigma and the strong silvery streak at the outer edge of the reniform spot, which is absent in M. confusa. The third related species, M. monosigna CHOU et Lu, 1979 has a reduced silvery pattern, the stigma is only a small spot and the streak of the reniform is absent. The configuration of the male genitalia also shows very special differences. Both confusa (Figs 3-4) and crassisigna (Figs 5-7) have symmetric, long saccular lobes and wide, ribbon-like basal cornutus in the vesica. (The differences between confusa and crassisigna are well-known, i. e. crassisigna has twice as many and much stronger cornuti in the vesica than in the case of confusa and the saccular appendage near to the ampulla is much longer). The new species has asymétrie, smaller saccular lobes, the valvae are more arcuate, the basal cornutus in the vesica is much more slender — but its sclerotization is stronger —„ armatures of cornuti show an intermediate form between the two species mentioned above; the shape of saccular appendages is similar to that of confusa. The third species, monosigna also has asymétrie saccular lobes, but these are longer, the basal cornutus of vesica, similarly to confusa and crassisigna, is wide. The female genitalia are very similar to those of confusa (Fig. 26) but the bursa copulatrix is constricted at middle stronger. On the basis of its morphological characters, the new species should be placed between confusa and crassisigna. These three species occur sympatrically in Korea, Japan, probably also in China and in the Amur-Region. Remarks — The new species was mentioned in a previous article as Macdunnoughia sp. (RONKAY 1982). Studies on the type of M. rhopalosema (HAMPSON 1913) have pointed out that the type specimen of rhopalosema is conspecific with the type of crassisigna, consequently the Korean species should be a distinct new one. It was found during investigations on the types of the presumed relatives that both of the two species mentioned previously as a members of the genus Macdunnoughia, namely camptosema (HAMPSON 1913) (Plate: 4-5) and schalisema (HAMPSON 1913) belong to the genus Autographa HÜBNER, 1821. The first species was already mentioned as Autographa by WILTSHIRE in 1971 without the note "stat. n.", the second is known only from the unique type, although Dufay described a new species from Burma {emmetra DUFAY, 1978) which may be synonymous with schalisema by as judged from the description and the photo. The genitalia of the types of these two species are figured on Figs 18-19, 30.