Kaszab Zoltán (szerk.): A Magyar Természettudományi Múzeum évkönyve 71. (Budapest 1979)

Szujkó-Lacza, J., Kováts, D. ; Rajczy, M.: Revision and numerical evaluation of the Hungarian Spergularia species (Caryophyllaceae)

Notes — We had to cite here the original description of this species pre- and ante-Linnean including LINNÉ (1762), because a group of authors believes LINNÉ described it in 1753, and others that only in 1762, an it is strongly discussed even today (cf. RAUSCHERT 1977). The first description of LINNÉ may refer to two or three species of Spergularia (S. rubra and S. salina), therefore, it is not valid, while the second (1762) contains some errors, or more exactly, insufficient description among the characters. The characters of Spergularia media were given first in an acceptable form by PRESL (1826). Spergularia rubra (L.) J. et C. PRESL (1819) — Syn.: Alsine Spergulae facie minor Bauch. Pin. 1st ed., 2:51, 1623 et 1671 — A. rubra Crantz Insl. 2:407, 1766;— Arenaria campestris All. Flora Pedem. 2:114, 1785. — A. rubra a campestris L. Sp. pi. 423, 1753 — A. (sect. Spergularia) rubra Pers. Syn. 1:504, 1805; Smalbg. cf. Gorschkova 1936; — Buda rubra Dum. Fl. Belg. 110, 1827; — Fasciculus ruber Dulac FI. Hautes-Pyrén. 245, 1867; — Lepigonum rubrum Whlbg. Fl. Gothoburg. 45, 1820-24; — Melargyra rubra Rafin. Fl. Teil. 3:81, 1836; — Spergularia campestris (L.) Aschers. Fl. Prov. Brandemb. 2:94, 1864; Hegi III. Fl. Mittel-Eur. 3:422, 1906; Aschers. — Graebner Syn. Mitteleur. Fl. 5:840, 1919; Groschkova Fl. URSS. 558, 1936; — S. subra a campestris Fenzl in Ledeb. Fl. Ross. 2:167, 1844; Gren and Godr. Fl. Fr. 1:275, 1848; Willk. and Lange Prodr. Fl. Hisp. 3:104, 1874; Fiori and Paoletti Fl. anal. Ital. 1:338, 1898; Gortani Fl. Friul. 245, 1867 — Stipularia rubra Haw. Syn. pl. succulent. 103, 1812; — Spergularia subra subsp. S. campestris Rouy and Foucaud Fl. Fr. 3:309, 1896; Briquet Prodr. Fl. Corse 1:489, 1910; — Tissa campestris Pax in: Engler-Prantl Nat. Pflanzenf.3. 1. b. 45, 1889. Though Spergularia rubra can also be found under different names it was characterized sufficiently by LINNÉ (1753). Later on, specialists, as LEBEL (1868), KINDBERG (1863) described several new species, compared with LINNÉ (1753, 1762), of the genus Spergularia (LEBEL I.e.), or of the genus Lepigonum (KINDBERG 1. c). Among these some new species resemble S. rubra, species differ from it in other characters like the surface of the seed. A number of specialists was confused by the fact that LINNÉ (1. c.) described S. rubra and a campestris also and the latter was characterized insufficiently. This was the source of an other mistake, and consequently numerous specialists used the name of "campestris" as a species name (ALLIONI 1785, ASCHERSON 1864, GORSCHKOVA 1936), or as a subsp. (ROUY & FOUCAUD 1896, BRIQUET 1910), sometimes it occurs in other genera as a species name (PAX 1889). Much later, besides LINNÉ'S description (I.e., p. 423): "Arenarie foliis filiformibus, stipulis membranaceis vaginantibus", PERSOON (1805, p. 504) characterized the species ".. .fol. filiformibus internodo brevioribus.

..". In 1819 these two descriptions were well combined by J. and C. PRESL, as Spergularia rubra. Spergularia salina J. et. C. PRESL (1819) — Syn.: Alsine atheniensis var. marina Sampaio Lista herb. Portug. 79, 1913; — A. marina Mert. and Koch Deutschi. Fl. 3:293, 1826; — A. marina a minor Koch Syn. Ill, 1835 — A. marginata C.A. Mey Verz. Pfl. Cauc. 217, 1831; — A. salina Groves in: Bab. Man. 9th ed. 681, 1904; — Arenaria Bocconi Soleiv. in: Stend. Nomencl. 2nd ed. 1:123, 125, 1840; — A. halophila Bge. in: Ledeb. Fl. Alt. 2:162-175, 1830; — A marina Pall. Reise. 3:603, 1766. Allioni Fl. Pedem. 2:114, 1785; Roth. Tent. Fl. Germ. 1:189, 1788;;— A. pubescens Stend. Nomencl. 1th ed. 66, 1821 ; — A. salina Ser. in: Dc. Prod. 1:401, 1824; — A. rubra ß marina L. Sp. pi. 1th ed. 423, 1753; — A. Spergula Dufour in: Ann gén. Sc. Phys. 7:293, 1820; — Buda media Dumort. Fl. Belg. 110, 1827; — Holosteum sperguloides Lehm, in: Ynd. sckol. 21, 1828; — Lepigonum caninum Leffl. OBZ. 19:103, 1869; — L. heterspemum Schur Enum. Fl. Frans. 126, 1866; — L. medium Wahlb. Fl. Gothob. 45, 1820-24; Fris Mant. 3:33, 1842. Koch Syn. Pond. 2nd. ed. 2. 121, 1843? — L. neglectum Kindb. Symb. Syn. Lepig. 6, 1856 — in: Lange Handb. Deutsche Fl. 3rd ed. 334, 1865;— L. neglectum et var. tenue Kindbg. Symb. Syn. Lepig. 6, 1856; — L. rubrum Wahlberg Fl. Gothob. 45, 1820; —

L. salinum Sweet hort. Brit. 3rded. 63, 1839; Kindberg Monogr. Lepig. 3-44. 1863; — Spergula filipaneis D. Dietor. Syn. pi. 2:1598, 1840; — Spergularia canina var. sperguloides Hartm. Handbok i scandinaviens 2nd ed. 248, 1832; — S. Dillenii Lebel in: Mém. Soc. sc. nat. Cherb. 14, 1869; Burnat Fl. Alpes-Marit. 1:273, 1892; Rouy and Foucaud Fl. Fr. 3:303, 1896; — S. halophila a marina Marss. Fl. Neuvorp. and Rügen 77, 1869; — S. heterosperma Heldr. var. sperguloides Fouc. in: Monogr. Illust. Spergular. m. in. Lebel 1868 — S. marina Barth, and Wendl. Beitr. Bot. 2:64, 1825; Griseb. Spie. Fl. Rum. Bith. 1:213, 1843; Willk, in: Willk. and Lange Prodr. Fl. Hisp. 3:165, 1880; Monniet-Ratter Fl. Eu. 1:155, 1964 — S. marina heterosperma Neilr. FI. Nieder-Österr. 782, 1859 — S. media Boiss. Fl. ov. 1:733, 1867 — S. media var. heterosperma Fenzl. in: Ledeb. Fl. Ross. 2:165-169, 1844-46 — S. media f. marina Clavaud Act. Sc. Linn. Bord. 35:402, 1881 — S. neglecta var ß salina Sowerby's lugl. Bot. 2:130-131, 1902 — S. rubra ß salina (1870) a germina ( 1874; Celak. ÖBZ. 20:48, 1870; Celak. Prodr. Fl. Böhm. 491, 1874 — S. rubra S marina Fiori and Paoletti Fl. anal. Ital. 139, 1898; App. 68; Gortani LI. Friul. 2:157,