Kaszab Zoltán (szerk.): A Magyar Természettudományi Múzeum évkönyve 70. (Budapest 1978)

Zombori, L.: New sawfly species in the Hungarian fauna (Hymenoptera, Symphyta), IV

First Hungarian locality: Hortobágy, Újszentmargita, védett erdő (nature conservation forest), fényre (collected at lamp), 1974. VI. 20, leg. GOZMÁNY. (HNHM). 4. Dolerus quadrinotatus BÍRÓ, 1884, sp. rev. — This species was described on the basis of two female specimens by the great Hungarian collector LAJOS BÍRÓ. A few years later KONOW (1891) treated the taxon only as a variety of D. thoracicus FALL. (p. 215). Ever since subsequent authors accepted this ranking, obviously, no one has taken the trouble to look at the type-specimens concerned. Since 1 am critically working myself through the sawfly material of the Hungarian museums, in Budapest, I carefully studied the type. I say type and not types, since there is only one type-specimen left in the collection, which was designated as "Lectotypus Ç Dolerus quadrinotatus BÍRÓ, 1884 des. ZOMBORI, 1977". The other type-specimen, it is most likely, is with DR. W. HELLÉN (Helsinki) who identified in the 1950s long series of sawflies from the Hungarian Natural History Museum, Budapest. Besides the type from Tasnád (today Tasnad, Transylvania, Roumania), several other specimens are known from a number of localities: Szőllőske (today Seleska, Czechoslovakia), Német Bogsán (today Bocsa Vasiovei, Roumania), Borosjenő (today Ineu, Roumania) and Simontornya (Hungary). As far as I am aware this species has not been recorded outside of the Carpathian Basin. The flight period of the 21 available specimens, all females, is between the 25th of March and the 14th of April, apparently an early species even among the representatives of the genus Dolerus. BIRO compared his D. quadrinotatus to D. vulneratus MOCSÁRY, 1878, described from Siberia, but BIRO'S species clearly differs from it in several characteristics. In order to find its place in the system of Dolerus the re-established quadrinotatus is included in a key for the species-group of thoracicus. This group of species belongs to BENSON'S (1952) so-called black-group of Dolerus with evenly punctured mesopleuron, black abdomen, 1st and 2nd abdominal tergites "always with some transverse surface sculpture", etc. This particular group may be separated from the rest of the black-group by their thorax being at least partly orange yellow (except the melanic race of sanguinicollis KL.) and their entirely black legs at least as far as the females are concerned. CONCISE KEY TO THE THORACIC US-GROUP OF SPECIES IN EUROPEAN DOLERUS (FEMALES) 1 (4) Metallic species, head and thorax with bluish black metallic reflection. 2 (3) Entire mesonotum, including front lobes black. Head parallel behind eyes haematodes (SCHRANK, 1781) 3 (2) Front lobes of mesonotum reddish yellow. Head somewhat contracted behind eyes rufotorquatus O. COSTA, 1864 4 (1) Non-metallic species, head and thorax without any metallic lustre. 5 (6) Pronotum entirely orange yellow (except in northern race) as are tegula, front and lateral lobes of mesonotum. Whole of mesopleuron black. Setae of sawsheath directed strongly laterally, then suddenly bending inward at apex sanguinicollis KLUG, 1814 6 (5) Pronotum entirely black; though front and lateral lobes of mesonotum may be marked with orange yellow. At least upper corner of mesopleuron also marked with orange yellow. Setae of sawsheath directed backwards, evenly arcuating over entire length. 7 (8) Front and lateral lobes of mesonotum entirely, more than half of mesopleuron orange yellow. Head and thorax with long pubescence, always much longer than diameter of front ocellus thoracicus (FALLÉN, 1808) 8 (7) Front lobes of mesonotum black, each with a small laterally situated orange yellow fleck, lateral lobes entirely black. Upper corner of mesopleuron marked with orange yellow extending to less than half of surface. Head and thorax with short pubescence, at most as long as (on mesopleuron) or much shorter (on head) than diameter of front ocellus quadrinotatus BÍRÓ, 1884