Kaszab Zoltán (szerk.): A Magyar Természettudományi Múzeum évkönyve 69. (Budapest 1977)

Hably, L. ; Csaba, A.: A new Smilax L. species from the upper Oligocene of Vértesszőlős (Hungary)

upper third of Smilax grandifolia (UNG) HEER is not of such a large extent as in S. tataensis: S. grandifolia of the Tertier has been obtained from several habitats (HEER 1855, STAUB 1864, KNOBLOCH 1958, GIVULESCU 1973). The form of Smilacites grandifolia UNG. (UNGER 1847) (Fig. 6) shows a certain similarity to S. tataensis, the venation are, however, entirely different. The primordiary veins accompanying the main vein embrace an ovate form, while the further veins run parallel with the margin of the leaf. As regards S. obtusifolia HEER (HEER 1855) (Fig. 3) even its shape is different, since the peak of S. obtusifolia is not an acute one, and the venation also shows a considerable difference which feature is the most essential one. In S. obtusifolia all the 7 veins take a course towards the apex, the marginal ones near the edge strongly hollow out and run parallel with the border of the leaf. ANDREANSZKY described two new Smilax species from Hungary (ANDREANSZKY 1956, 1959): S. borsodense (Fig. 5.) and S. praeaspera (Fig. 4.), both considerably differing from S. tataensis. The new species was recovered in two specimens from the flora of Vértesszőlős. The paratype is with a counterproof. The length of the latter is 9.0 cm, its width is 6.2 cm in the upper third of the blade. The course of the right-side vein alongside the main vein is 43°, that of the outer vein is 124/84°. A steeper course is characteristic of the left-side veins. According to Table 1. Smilax tataensis show dissimilarities from the other four species in two exomorphic characters: 10 and 13; and in three quantitative characters : 3, 4 and 5. As we can seen in the Table 2. common for the every earlier and here described species is the character number 14, therefore it is a generic character up to date. In the character number 1 and 6 four species show similarities, the character numbers 9, 11 and 12 are common in three species, while character numbers 2 and 8 are common only in two species. References ANDREANSZKY, G. (1956): Neue und interessante tertiäre Pflanzenarten aus Ungarn. II. — Ann. Hist.-nat. Mus. Nat. Hung., 7 : 221-229. ANDREANSZKY, G. (1959): Die Flora der Sarmatischen Stufe in Ungarn. — Budapest, Akad. Kiadó, p. 182, Abb. 234, 235, 236. GIVULESCU, R. (1973): Ein Beitrage zur Kenntnis der fossilen Flora des SchiltalS. — Mémoires (Bucuresti), 19: 23. HABLY, L. (1976): A vérteSSzőlősi Baromállás felsőoligocén korú flórája. —Szakdolgozat. HEER, O. (1855): Flora tert. Helv. I. p. 82. T. XXX. fig. 8. KNOBLOCH, E. (1958): Die oberoligozän Flora des Pirskenberges bei Slukov in NordBöhmen. — UUG. Sbornik 16. XXVI., paleontologicky, p. 251. Taf. XIII. Fig. 7; Taf. XV. Fig. 9. STAUB, M. (1887): A Zsilvölgy aquitánkorú flórája — M. Kir. Földt. Int. Évk. 7: 251, T. XX-XXI. fig. 1-7; XXII-XXIII, fig. 1-5; XXIV. fig. 1; XXXIV-XXXV. fig. 1 c . UNGER, F. (1847) : Chlor, prot. p. 129. T. XL. fig. 3. Address of authors: LILLA HABLY Botanical Department Hungarian Natural History Museum H-1146 Budapest, Vajdahunyadvár Hungary ATTILA CSABA Dózsa György út 12 H-2027 Dömös Hungary