Kaszab Zoltán (szerk.): A Magyar Természettudományi Múzeum évkönyve 66. (Budapest 1974)

Ujhelyi, J.: Data to the systematics of the sections Bulbosae and Caespitosae of the genus Koeleria, XII.

lia, glabrescentia, vel sparse pilosa, auriculata, vaginis usque 4 cm longis, glabriusculis in fauce ciliatis. Ligulae rotundatae, 0,8 mm longae. Gulmi plantarum evolidarum pro more supra mediam foliati, laminae foliorum culmeorum usque 6 cm longae, et •0,7 mm latae, acutae, apice semper convolutae, glaberrimae tantum scabriusculae, auriculatae, vaginis usque 9 cm longis, infimis br éviter pubescentibus mediis superioribusque glaberrimis vel tantum glabriusculis. Ligulae longiores, rotundatae, cca 1,5 mm longae, ad marginem crenatae. Gaules usque ad 28 cm longi et 1 mm crassi, glabri. Paniculae usque 10 cm longae et 1 cm crassae, subinterruptae, lobatae, laxiores, oblongo-cylindricae, albescentes sed tantum subnitentes, sericeae. Spiculae parvae, 3,5—4 (4,5 mm) longae, et 1,5 mm latae, biflorae, superficie minutissime parce hirsutae. Glumae inaequales, inferiores 3,5 mm longae, lanceolatae, superiores 4 mm longae, ovatae, acuminatae ; lemmae 4 mm longae, acutae, superficie minute puberulae, paleae 3,5 mm longae, bicarinatae, hylinae, acutae. Antherae 2 mm longae. (Plate I Fig. 5) Gellulae epidermidis laminae foliorum innovationum parvae. Series cellularum costalium 1—4, intercostalium 5—10. Habitat in graminosis Vallesiae, Sueciae. Typus meus: Koeleria alpigena m. Brançon (Vallesia) 1. VI. 05., A. MAILEFER H. L. <Dupl. in herb. Domini) (PRC). Description of the leaf epiderm: On the abaxial and axial sides of the innovational leaves the cells are small and comparatively short. Senile leaves with weakly developed costal zone, occasionally even absent, 1—3-seriate, (in contrast to every other Koeleria species treated here to fore there are also hairs in the costal zone, while the large parenchymatous cells exceed by their dark color when stained, and by the application of the Vezuvin-malachite double staining they turn dark red, indicating a pure cellulose cellular wall which, at the same time, are also heavily incrassate. I found this phenomenon only in some other tribes of the family Qramineae. Furthermore, sclerenchymatous bundles extend also below such fields where the epidermal sturcture is identical with that of the epidermal zone. This phenomenon appears in the species of the Series Pubiculmes UJH., from the North Carpathians and the Austrian Burgenland, and in Koeleria filifolia (DOM.) UJH. from the Spanish Sierra Nevada. Under the epiderm of such blades there developed contiguous sclerenchyme, but in the species under discussion this is not regular)

. Costal zone with the relatively short parenchymatous cells irregularly alternating with sporadically appearing oblong silica-bodies. Between them, parenchymatous cells alternating with primordia of minute macrohairs. The stronger, therefore 3-seriate costal zone reveal bigger macrohairs, and also more silicabodies. (According to the general experience, no macrohairs grow in the costal zone. In the present case the sclerenchymatous fibres of the costal zone are undeveloped, and in the uniseriate zone the oblong silica-bodies frequently alternate with heavily developed parenchymatous cells). Intercostal zone, where costal zone present, 10—11 seriate, the parenchymatous cells regularly alternating with primordia of minute macrohairs. Occasionally some parenchymatous cells strongly staining in middle of zone, therefore of thick walls; stomatal cells appearing in one row each on both sides of zone, alternating with elongate parenchymatous cells ; these rows without primordia of. small macrohairs. Costal zone of juvenile blades 2—4-seriate, well developed, parenchymatous cells narrow, alternating regularly with oblong or cubicoid silica-bodies, their walls weakly sinuous, thin. Parenchymatous cells only rarely (in one or another place or the leaf) replaced b}^ primordia of small macrohairs, sporadically showing also cork-cells. Intercostal zone 6—8-