Kaszab Zoltán (szerk.): A Magyar Természettudományi Múzeum évkönyve 65. (Budapest 1973)
Ujhelyi, J.: Data to the systematics of the sections Bulbosae and Caespitosae of the genus Koeleria, XI.
primordia of minute micro-hairs, with only occasionally appearing one or two bigger micro-hairs primordia ; occasionally a costal zone omitted, then field 9-seriate, medially usually with 2 rows of stomatal cells, in general alternating with a parenchymatous cell, and rarely with an inserted microhair primordium (Plate III, Fig. 1-2). The medium high, about 50 cm tall plant is densely cespitose, owing to intravaginal innovation some sterile shoots strongly appressed to one another; shootgroups originating from the short, decumbent, 2 cm long rhizomes, w T ith one flowering culm each, rhizome with short internodes, densely rooty. Vetust vaginae whole or only very slightly lacerate, pale yellow, only 1.7 cm long in type specimen, sericeous with very minute and appressed pubescence. Senile leaves short, merely 4-5 cm long, 1.5 mm wide, weakly explanate or flat. Blades glabrescent, marginally with very small prickle-hairs, hence very finely puberulous. Leaf base slightly auriculate, with a weak carnose incrassation. Ligulae minute, only 0.5 mm long, abrupt, marginally ciholate. Juvenile leaves slightly longer, reaching even 6 cm, narrow, at most 1 mm vide, convolute, pale grayish-green, midrib and margins puberulous owing to presence of minute prickle-hairs. Vaginae about 2 cm long, slightly longer than those of senile blades, their hgulae bilobate and marginally ciliolate. Developed culm foliose to about one-third of its length; blade of uppermost leaf about 3.5 cm long, apically attenuate, about 2 mm wide, glabrous. Ligulae 0.5 mm long, bilobate, apically rounded, marginally also ciholate. Culm 34-39 cm long, glabrous, 1 mm thick. Panicle oval — cylindrical, 2.5-4.5 cm long, nearly 1 cm thick, compact, in its inferior part slightly lobate, pale violaceous (variegated), hirsute. Spikelets 0 mm long, 2.5 mm wide, ovate, biflorous, surface rarely pubescent. Glumae of inequal size, lower one 4.5 mm long, lanceolate, upper one 5.5 mm long, apically attenuating, rarely hirsute w T ith long hairs, its carina scabriusculous. Lemmae 5.5 mm long, aristately attenuating, their surface minutely puberulous. Paleae bicarinate, hyaline, apically obtuse. Antherae 2.8 mm long. The plant, regarded as Koeleria splendens PRESL (see Achtarov: Koeleria splendens PRESL var. pyrinica ACHT., from his herbarium, ACHTAROV 1951, JORPANOV 1963) owing to its considerably smaller stature, gracile culm, narrow panicle, bulbosely incrassate rhizome, convolute, short and narrow^ blades, differs at the first glance from PANCIÖ'S type. Whereas Koeleria tuzsonii UJH. has a habit indicating a more xerothermous habitat, and thus its stature resembles that of Koeleria penzesii UJH. (which also has panicles with a violet suffusion), its descendant, the polyploid Koeleria eriostachya PANÖ. is a decidedly robust plant with a strong and large rhizome, therefore loosely and elongately cespitose, and with a large violet panicle, in short an expressedly alpine to subalpine life-form plant. Owing to these very characteristics, it was connected with the also subab,tinealpine Koeleria carniolica KERN., and drawn in this latter as a sysnonym. NOVAK'S specimen from the Rila, the same as PANCIC's plants from Gobelja, unmistakably bear aU morphological and anatomical features of Koeleria eriostachya PANÖ., despite the fact that the differences are not only those of dimension. The plant assumably lives not only in the localities listed, but it failed to draw the attention of collectors, unlike its taller, strongly pubescent descendant, Koeleria eriostachya PANÖ. If DOMIN had seen NOVAK'S specimen, or if he did see it in some collection which escaped my attention, he would have considered it some variety of Koeleria splendens PRESL, owing to its bulbosely incrassate base and convolute leaves. ACHTAROV was of the same opinion. Only a more thorough study, the hirsuteness of the culm below the panicle and the pubescence of the panicle reveal that
