Kaszab Zoltán (szerk.): A Magyar Természettudományi Múzeum évkönyve 65. (Budapest 1973)

Ujhelyi, J.: Data to the systematics of the sections Bulbosae and Caespitosae of the genus Koeleria, XI.

DOMIN placed the area of Subsp. I. K. eriostachya PANCIC to the boreal, alpine* and subalpine meadows, pastures, and montane rocky sites of Bulgaria, Serbia, Croatia, Carniolia, Carinthia, Styria, Tirolia, Vallesia and Italia borealis. 1. var. typica DOM. represents specimens with wholly to largely villose to hirsute spikelets. Excepting Vallesia, he published exemplars from the entire area of the subspecies. 2. var. carniolica (KERN.) DOMIN occurs, according to DOMIN, in Serbia meridionalis, Croatia, Carniolia, Littorale, Carinthia, Styria, Tyrolia and Vallesia. This taxon differs from the typical by the indûment of the spikelets being slighter, and with their carinae more or less weakly ciliate or indeed glabrescent. 3. var. engadensis DOMIN inhabits the Engadin in Vallesia. The main characteristics of Subspec. ILA". Schroeteriana DOM. are given as follows : the usually wholly pubescent culm is gracile, lower, the blades narrower, the vaginae pubescent, the panicle less coloured, or pallid, the spikelets merely 5 mm long, the glumes shghtly shorter. Its area covers the montane rocky sites, the stony swards and pastures of the subalpine and alpine regions of West Vallesia, East and Southern Gallia, as well as the Pyranó:^. I should like to emphasize already here that aU these plants belong to the Series Arenariae ÚJHELYI. I shall not discuss the two other Asiatic subspecies in the present paper. In Subspecies I. K. eriostachya PANC., the members of two evolutional series, sharply differing in area, evolution, morphological and anatomical structure, can be recognized. The specimens originating from the Canton Graubünden are identical with Koeleria lamarckii ÚJHELYI. Whereas the undersides of the leaves of the East Balkanian Series are invariably regularly hirsute, and the culm always hirsute or pubèrulous under the panicle, in plants from the SE Alps there appear at most only here and there some irregularly spaced minute micro-hairs on the undersides of the blades, and the panicle is only occasionally pubescent (DOMIN relegated these specimens to 1. var. typica DOM.), but usually glabrescent. Nor is the culm always densely pubescent under the panicle. DOMIN contracted the species to be discussed in Subsp. I. eriostachya PANC. owing to the similarity of their external apperance: their robust stature, ample and long innovations, compact, large and hirsute panicles, and greenish leaves, and to literature having widely drawn together the two species. The SE Alpian Koeleria carniolica KERN, was invariably mentioned as the synonym of the East Balkanian Koeleria eriostachya PANÖ. (HACKEL 1879, VELENOVSKY 1891; ADAMOVIC 1904). He could not set himself free from this general view. It should not be forgotten that we are deahng with polyploid plants which all live in subalpine to alpine regions, therefore in closely related ecological conditions. Also these species, like the members of the great-species Koeleria splendens PRESL or those of the collective species Koeleria pyramidata, are convergent species born for highly similar sites! XVII. Series Tuzsoniae ÚJHELYI ser. n. Rhizomatibus densis, culmis robustis, infra paniculam hirsutis, foliis plants, virescentibus, vel parce glaucescentibus, hirsutis, saepe denticulatis, vaginis minute adpresse hirsutis, paniculis pro more coloratis, subdensis, spiculis acuminatis, vel acidis, hirsutis.