Kaszab Zoltán (szerk.): A Magyar Természettudományi Múzeum évkönyve 65. (Budapest 1973)
Jánossy, D.: New species of Episoriculus from the Middle Pliocene of Osztramos (North Hungary)
Episoriculus borsodensis sp. n. Derivatio nomini s: borsodensis, named after the administrative department Borsod with the locality Osztramos. T vpe locality: Loc. 1, Osztramos Mill, Ne — Hungary (48° 30'North. Lat., 38" 25' East Long.) Type level: Middle Pliocene, horizon of Estramontium (see Jánossy, 1972). H o I o t y p e: Left maxillary fragment with the penultimate and last unicuspids and P 4 . Inventary number: V. 72. 115, Coll. Pal. Dep., Nat. Hist. Mus., Budapest. — Paratype: Anterior fragment of left mandible with the incisor, two antemolars and M 1 (see JANOSSY 1972, Plate II., Fig. 3). Referred specimens : Nearly a hundred very fragmentary remains, all representing teeth and diverse mandibular parts. Diagnosis : A small Episoriculus species, with relatively reduced anterior dentition (chiefly upper and lower incisors), as in E. tornensis sp. n., but last upper unicuspid and first lower antemolar larger. Ratio of P 4 /AP~'~ and the articular facets of mandible differing from that of all hitherto known fossil species. Description : I submitted a detailed sketch of the features of the genus Episoriculus as well as the comparison of E. gibberodon , E. tornensis sp. n. and E. borsodensis sp. n. in the course of the description of the former species. Suffice it here therefore to emphasize only the most important features of the new species of Locality 1, Osztramos. 1 described these remains as ,,? Episoriculus sp." on the basis of a very scant material originating from the locality mentioned (JÁNOSSY 1972). Among the various maxillary fragments coUected earlier and in the latest material of excavations in 1972, I found one specimen with a well preserved last upper unicuspid and two former specimens in which this tooth is represented only by its alveolar holes (see Plate I, Fig. 6). I became convinced that the form from Locality 1, Osztramos, is in this feature clearly betw r een E. tornensis and E. gibberodon . Beyond the diagnistic features, I have to emphasize the presence of quadratic (and not compressed) upper unicuspids, the less reduced first lower antemolar (against E. tornensis) and the narrower lower molars — all characteristics analyzed in detail in the course of the description of E. tornensis sp. n. given above. Pigmentation — if any — is the same as in E. tornensis: a light orange colour only on the top of the incisors and on the most prominent cusps of the last upper premolar and molars as well. A comparison with recent species as well as with the insular fossil form Nesiotites was also made above. The remains are comparable stratigraphically and morphologically for European fossil material, chiefly with the Podlesice-form (see KOWALSKI 195(i. Plate II Fig. 3). This figure is for a comparison somewhat too small, but it still seems to show the brachygnathous form, which appears to agree with E. borsodensis sp. n. Not only the forms from Beremend and Villány, but also the specimens originating from Csarnóta and Weze (by courtesy of M. KRETZOI, I received these latter from his collection) correspond with the elongated "dolichognathous" lower anterior dentition of E. gibberodon. Going by MEIN'S exact figures of the recently published remains from Rhodos the characteristics are again the same, in spite of the smaller dimensions (MEIN in: BRUIJN, DAWSON & MEIN 1970, Plate 11. Fig. lc). As mentioned above one can hardly form a valid opinion of the evolutionary connection between E. gibberodon, E. tornensis and E. borsodensis. Against our hy-
