Kaszab Zoltán (szerk.): A Magyar Természettudományi Múzeum évkönyve 64. (Budapest 1972)

Jánossy, D.: Middle Pliocene microvertebrate fauna from the Osztramos Loc. 1. (Northern Hungary)

proposed the establishment of a separate subfamily Amblycoptinae, but this was not accepted in later literature and our genus is now placed in the subfamily Soricinae and the tribe Neomyini REPENNING, 1967. A comparison with the otherwise very nearly related form ,,Anourosorex" kormosi BACHMAYER & WILSON, 1970, from the Lower Pliocene of Kohfidisch (Gyepüfüzes), would be meaningless, because this species possess 3/3 molars (M

3 is present). This form, which may be an ancestor of Amblycoptus and an extinct genus, is a much less specialized form than the remains from Hungary. Amblycoptus topali may be an end form or a nearly end form of a fine evolutionary lineage of these peculiar shrews. Eomyidae Leptodontomys SHOTWELL, 1956 Leptodontomys bodvanus n. sp. Derivatio nominis: „bodvanus" from the river Bódva in the valley bordering the hill Osztramos in which the material was washed. Stratum locusque typicus: Osztramos, locality 1. (further particulars see at the description of Amblycoptus topali). Age: Middle Pliocene („Postpannonian Pliocene"). Diagnosis: A small Eomyid from the Eomys-Leptodontomys group, near Leptodontomys {„Eomys") catalaunicus HARTENBERGER, 1966, differing from it by the weaker posterolophid in the lower molars (Mi

/o

) and the stronger posterolophs in the upper molars (M

1

/*). Holotype : Right mandible with the first and second lower molars (Mj and M

2

), the alveolus of M

3 and the partially broken alveoli of P

4

; lacking the ascending ramus and the anterior part of the mandible (see Plate III. fig. 1); No. V. 71. 52 in the Natural History Museum, Budapest, Hungary. Pa rat y pes: Slightly worn right upper M

1

'*. Referred specimens: subadult and strongly worn M

1

^ and Mi/, (see Plate III. Pig. 2-5). Description : Both the upper and lower cheek teeth are distinctly brachyodont, similarly as in the different species of the Eomys (s.str.) - Leptodontomys evolutionary line. The crown of the upper first and second molars consists of a protocone-paracone and metacone-hypocone ridge, connected with a longitudinal crest. The mesoloph is reduced, the anterior cingulum T-shaped, but lingually reduced. The posterior cingulum connects to the hypocone in the same way as in cricetids, it is much more strongly built as in L. catalaunicus (with an accurate labial angle). The anterior and posterior cingula lie deeper than the main cusps. The premolar and third upper molar are lacking. The structure of the lower first and second molars is nearly the same as that of the upper ones. The anterior cingulum is separated in the young specimens. In the process of abrasion, this cingulum becomes connected at the center of the tooth with the protoconid-metaconid ridge and extends both lingually and labially forming a T. There is either a reduced mesolophid, or a stronger one extending lingually in a weak spur. The posterior cingulum is, in contrast to L. catalaunicus, very weak, connected to the hypoconid only.