Kaszab Zoltán (szerk.): A Magyar Természettudományi Múzeum évkönyve 64. (Budapest 1972)
Ujhelyi, J.: Data to the systematics of the sections Bulbosae and Caespitosae of the genus Koeleria, X.
in Leclb. Fl. ross. IV., p. 401 (1853) pro syn. K. cristatae p. p. — Koeleria cristata b major GBISB. in Ledb. Fl. ross. IV. p. 402 (1853) p. p. — Koeleria cristata var. villosa;MEINS Herb. Fl. ingr., cent. X. No. 772 Bp. (1874). — Koeleria cristata var. dasyantha TRAUTV. Herb; Koeleria polonica DON. Magy. Bot. Lap. III. p. 180 (1904) ampl. var. typica DOM., 1. e., p. 244. Prov. Leningrad , distr. Luga. In pineto sicco pr. st. viae Ferreae Tolmaczevo. Leg. S. GANESCHIN. 192(3. VII. 14. (BP). — Koeleria polonica DOM. (an Koeleria grandis BESSER?) Janów pr. Leopolim (Galiciae) — in silva caedua "Jarina" dicta, solo calcareoarenoso, frequens., 16. VI. 1909. Legit Hof. BLOCKT (BP) ; Hungária: J. de Hulják, Plantae Hungáriáé exsiecatae. Koeleria gracilis PERS. Nyírség. A község elején, az útmenti száraz tölgyes-erdőben. Mintegv: 140 t. sz. f. m. Szatmár: Bátorliget, 1935. VI. 9. Hulják. No. 219713 (BPU.). The plant is robust, loosely eespitose. Similarly to its progenitor, the lose eespitosity derives from the extravaginal innovation, resulting in the partly running and ramifying rhizome. Stature high, attaining even 90 cm (Reichenbach "'Flora Germ. exs."). but usually 00-70 cm high. Sheaths of basal leaves densely hirsute, villose, pili 1.2 mm long. Old sheaths whole, rugose, or emorse. Rhizome 7-(9) cm long, 1.2 mm wide, ramifying. Senile blades 10-20 cm long, 3 mm wide, green, weakly auriculate, flat, with a sericeous sheen owing to erect, long hairs. Their sheaths 5 cm long, ligulae abrupt, crenately margined, 0.8 mm long, ciliolate. Juvenile blades 8-11 cm long, 1.8 mm wide, glabrescent. with 4.5 cm long sheaths. Ligulae also abrupt, 0.2 mm long. Developed culm foliose to half its length, 2-3-foliate, blades 6-9 cm long and 3 mm wide, apically acuminate; sheath long, 18—21 cm. Ligulae 1 mm long, crenate, ciliolate. Wholly developed culm attaining 90 cm in height, but generally 65-70 cm long and 2 mm wide, in its upper part densely pubescent, with 0.5 mm long cilia. Panicle robust, loosely lobate, peduncle of lower lobe attaining 2 cm length. In its lower part frequently interrupted, 7-16 cm long and 1-2 cm wide, with hirsute rachi and ramuli. Spiculae 7 mm long, sometimes longer, biflorous, with a sericeous surface, occasionally appressedly hirsute. Glumae unequal, lower one generally 4.5-, occasionally 5 mm long, lanceolate, acute, upper one 6-, occasionally 7 mm long, elongately ovate, carina finely scarious. Lemma 5.5-6 mm long, lanceolate, sericeous, apically acute, palea 5.2-5.5 mm long, bidentate, hyaline. Anthera 2 mm long. By this exposition of the evolutionary trend, I hope to have correctly clarified these East European species of peculiar cytology. It is to be hoped that the other members of the series will also be discovered. The Series was doubtless born in a colder era, as substantiated by its area, and then extending to the shores of the Baltic Sea during a warmer period. Its origin in colder times is supported by its occurrence as a relict plant, among other relict taxa of the glacials, in the Natural Reservation Bátorliget, Hungary (co-existing there with Ligidaria sibirica (L.) CASS. Trollius europens L., etc.). Unfortunately, J. HULJÁK failed to recognize the plant as new, having identified it as the common Koeleria gracilis PERS., as known at that time. It is rather striking that no one else had collected this plant, in the well explored region of the Nyírség: a valuable find for the entire Carpathian Basin. B\ r his discovery, the flora of Hungary increased by a priceless taxon also from the point of view of floral history.
