Szekessy Vilmos (szerk.): A Magyar Természettudományi Múzeum évkönyve 62. (Budapest 1970)

Ujhelyi, J.: Data to the systematics of the sections Bulbosae and Caespitosae of the genus Koeleria. IX.

have a glabrous or but very finely hirsute panicle and leaves whose underside are evenly covered with erect pili. It was among these specimens that I found the hexaploid grade members of Koeleria eristata (L.) PERS. em. BORB., and Koeleria majorijlora (BORB.) BORB. The holotype of the plant described under the name Koeleria mollis MANN belongs also here. Both the morphological and anatomical structure of this species reflect those of the progenitors mentioned above. A part of Koeleria pyramidata (LAM.) DOM. var. villosa DOM. is assignable to Koeleria tristis DOM., the member of a distinct Series. The taxa of the alpicolous Koeleria pyramidata (LAM.) DOM. are the higher polyploid grade members of another independent evolutional line, namely of the Series Ciliatae ÚJHELYI. Scries XIV. Molles ÚJHELYI, scries nova Vaginis puberulis, foliis pubescentibus hirsutis et longe cilialis, paniculis valde elongatis, oblongo-pyramidalibus, vel pyramidalibus. The species belonging to this Series had been listed under the names Koeleria gracilis PERS., Koeleria pyramidata (LAM.) DOM., Koeleria pseudocristata DOM., and Koeleria cristata auct. I have already pointed out in my previous Koeleria papers (8. 9, 10) that the taxon Koeleria gracilis PERS. is a nomen dubium. Accordingly, and following BORBÁS'S excellent realization of the facts, 1 published (8) the diploid grade member of the Series as Koeleria cristata (L.) PERS. em. BORB., and the tetraploid grade member as Koeleria majorijlora (BORB.) BORB. I designated the series, of which at that time I knew but two members, temporarily under the name Series Cristatae. However, this name may cause misunderstandings owing to its use by K. DOMIN. It was also evident that I had to search for the higher polyploid grade members of the Series in Koeleria pyramidata (LAM.) DOM., and Koeleria pseudocristata DOM. Ulis genetically indubitably interrelated evolutional series is characterized by a denser or looser cespitose growth, wider and explanate basal leaves with only the juvenile ones being convolute (especially in drier habitats). In both the upper and lower sides of the blades, the parenchymal cells alternate regularly with considerably developed pili, beside the minute macrohairs, in the intercostal zone, so that the leaves are sparsely hirsute or villose. In stature and conformation, they stand nearest to the species of the Series Arenariae UJH., but whereas these latter are evenly hirsute owing to the development of the small macrohairs, the leaves of the species belonging to the Series Molles UJH. also possess the pili. The members of the Series Arenariae are thus not villosely but sericeously hirsute. Besides, their culm is also sericeous. The blades of the members constituting the Series Transsylvanicae UJH. lack the pili, and even the small macrohairs appear but sporadically; they are also glaucous. The leaves of the species relegated to the Series Pseudoglaucae UJH. are glabrous, with, occasionally, only initial macrohairs. The authors concerned, including also DOMIN, treated Koeleria mollis MANN as a high polyploid grade member of the Series Ciliatae UJH., as a synonym of Koeleria lamarckii UJH. = Koeleria pyramidata (LAM.) DOM. The leaves of the grasses assigned to the Series Ciliatae UJH., are invariably glabrous, with only their margins filiate. On the other hand, the panicles of the members relegated to the Series Molles UJH. are acute or acuminate, glabrous or sericeous, occasionally very finely puberulous. By the increasing polyploidy, the rachi of the panicle elongate (the HÍ*