Szekessy Vilmos (szerk.): A Magyar Természettudományi Múzeum évkönyve 58. (Budapest 1966)

Ujhelyi, J.: Data to the systematics of the sections Bulbosae and Caespitosae of the genus Koeleria. VI.

specific identity of the two taxa. For my part, since I had hitherto seen only E. HACKEL'S specimen, I, too was unaware of their identity, but now I saw its distinctness, differing from Koeleria caudata (LINK) STEUDEL. On the basis of the description, I presumed that Koeleria dasyphylla WILLK. is a derivation of an even higher ploid grade. The specimens preserved in the Madrid Museum from the Sierra Nevada, the •Sierra de las Nieves, and the Sierra del Pinar-Grazalma, agree both in habit and in the structure of every organ. The holotype of Koeleria nevadensis (HACK.) UJH., collected by HACKEL, differs only in its habit, the same as M. GANDOGER'S exemplar from Cantabria (the holotype of Koeleria caudata STEUD. var. pubiflora DOMIN). The culm of both specimens is shorter, the rhizome short, woody, the rib of the glumae ciliate. The habit of these two latter exemplars refer to a site of unfavourable substrate conditions. The plant is cespitose, its rhizome elongated, generally 3 cm long, and slightly stolonate due to the recumbent rhizome. The rhizome is covered with the large, entire sheaths of the vetust senile leaves. The recumbent, loose rhizome refers to also a looser lawn, in contrast to the two preceding species. Generally, the plant is 35—45 cm high, with 3—4 blades on the culm to about its half height, the highest leaf is 2 cm, its vagina 6 cm long, the culm about 1 mm thick, basally recumbent. The senile leaves are invariably flat, 2—4 cm long, about 2 mm wide; the juvenile blades might attain 6 cm, they are narrower, plane or convolute; both kinds of leaves are densely velutino-pubescent, together with the entire sheaths, marginally with scattered, erect, large cilia. The ligule is about 1 mm long, marginally crenate, ciliolate. The panicle is 4—5 cm long and 1.3 mm wide, ovally cylindrical, looser than in all preceding species due to the slightly stalked spiculae, these latter 5—6 mm long, glabrous, greenish, elongate. The inferior glume is 4, the superior glume 6 mm long, the lemma 4, and the palea 3 mm long, all of them aristulate. The anthera exceed even 2.2 mm (Plate I, Fig. 6). Specimina examinata: In rupestribus glareosisque (Loc. class). Sierra de las Nieves (Málaga), 11, VII. 1930, C. Vicioso, sub nomine Koeleria dasyphylla WILLK. (MA, nr. 9810) ; Sierra del Pinar-Grazelma (Cadiz), in saxosis, 15, VII. 1930, CEBALLOS et VICIOSO, sub nomine Koeleria dasyphylla WILLK. (MA, nr. 9811) ; In saxosis glareosisque schistosis (Los. class.) Sierra Nevada; vertiente oriental del Cerro Caballo (Granada) 2.900 m.s.m., CEBALLOS et Vicioso, 8. VII. 1930, sub nomine Koeleria caudata (LINK) STEUD. var. nevadensis HACK. (MA, nr. 9772). Description of the leaf epiderm : On the abaxial side of the innovational leaves the epidermal structure is as follows: In the senile leaves the costal zone is 2—3seriate, the cells of the basic tissue are elongate, wider than in the preceding species, straightly decurrent, very finely sinuous, alternating with elongate cilia, macrohairs and sporadically with large, cubic silica-bodies; the intercostal zone is 4—5seriate, with 1—2 stomatic rows, stomata bigger than in preceding species, the cells of the basic tissue are strongly elongated, alternating with cilia, the intercostal zone with 1 stomatic row, the longitudinal wall of the cells of the basic tissue is slightly sinuous, the cellular walls minutely sinuous. The structure of the epidermis and the measurements of the cells refer to a hexaploid plant (Table IV, Fig. 1). Koeleria dasyphylla (LINK) STEUDEL var. nevadensis (HACK.) ÚJHELYI combinatio nova Plantae dense caespitosae, folia vaginaeque vestitae, vel scabrae, ad mar ginem ciliatae. Glumae et lemmae in carina ciliatae, caeterum sparse hirsutae (Tab. II. Fig. 8.). Habitat in Sierra Nevada et in Cantabria.