Boros István (szerk.): A Magyar Természettudományi Múzeum évkönyve 7. (Budapest 1956)

Baksay, L.: Cytotaxonomical studies on the Flora of Hungary

shores of the Danube. The chromosome numbers agree with the ones of the other related species examined in Europe. 33. ! Physocaulis nodosus (L.) Tausch. 2n = 22 (2x). Lábatlan, Mts. Gerecse, Pisznice ; Querceto-Cotinetum, on limestone. This annual species is the sole representative of the genus. Its range in the Mediterranean extends from Portugal to the Crimean, the Caucasus and Iran. It is an ancient Mediterranean, (tercier?) relict, species in Hungary. 34. ! Bupleurum falcatum L. 2n = 16 (2x). The Solymár wall ; FestucoBrometum erecti, on dolomite. — Bánkút, Ablakoskö valley, Mts. Bükk (900 m) ; Seslerio-Fagetum bükkense, on limestone. 35. ! Bupleurum dilatatum Schur, herb. Trans. 2n = 32 (4x) (B. falcatum L. var. d. latifolium Schur Sert. n. 1162 ; B. falcatum ssp. dilatatum Schur En. pl. Transs. ; ad B. petiolare (?) Lapeyr. Hist. Arb. pl. Pyren ; et aut. plur., vide : Wolff in Engl. Pflanzenreich 43. IV. 228). Mt. Csókakő, Mts. Vértes ; Fageto-Ornetum, on dolomite. The B. falcatum L. s. 1. conspecies is a highly variable and a very extensively ranging hillymontane Eurasian species. It has a very broad oecology, and is a member of extremely different plant associations. It is frequent in Hungary in relict habitats, on grassy slopes, further also in woodlands. The chromosome basic number of the genus is 8 and there was no polyploid known among them. The above data explains the cause of the variability, on the ground of which the whole specific complex will have to be revaluated. The polyploid species was possible to show from numerous points in Europe, on the basis of comparative morphological and epidermis examinations made on herbaria material. The plant examined by the author agrees with the cotype of Schur. The exact range and oecological claims of this species require further studies. 36. ! Ferula sadleriana Ledeb. n == 11 (2x). Mt. Pilishegy, Pilisszántó ; Querceto-Cotinetum. An endemic species in the Carpathians' Basin, — a praeglacial relict. Its chromosome numbers agree with those of the species examined hitherto in the genus. 37. Asperula taurina L. 2n = 22 (2x). Mt. Zengővár, Pécsvárad, Mts. Mecsek ; Fagetum. Leg. Újhelyi. A Mediterranean species, occurring only in the southern part of the country. 38. Knautia arvensis (L.) Coult. 2n = 20 (2x) Szár, alongside roads, on loess. — Alsógöd, on sand ; Festucetum sulcatae. — Mt. Bálvány (900 m) Bánkút, Mts. Bükk ; Festucetum sulcatae. A highly variable species, both on lowlands and in mountains, in the whole of Europe. 39. ! Knautia Kitaibelii (Schultes) Borb.* ssp. tomentella (Szabó) Baksay. 2n = 40 (4x). Mt. Nagyszénás, Pilisszentiván ; Fageto-Ornetum, on dolomite. This species belongs, with reference to its morphological characteristics and especially its synpodially branching rhizome, to the subsection Silvaticae in the section Eutricherae of the genus Knautia, and not into the subsection Arvenses wherein Z. Szabó had relegated it. Into this latter subsection belong species with taproots, wherein the shoots are situated in the level of the soil so that the upper part of the root is branching (multiceps). Kn. Kitaibelii ssp. tomentella breeds, according to my observations, vegetatively, supported also by A. Pénzes (verbal communication). It is an expressedly umbrophilus species : planted in the sun in the Botanical Gardens it does not bloom, which character together with its slender form also connects it up with the Silvaticae subsection. The main species is a subalpine, Carpathian endemism, segregated already by V. Borbás under the name „A). praealpinae" from the group „B) Arvenses". The working out of the race groups of the species will be the object of another paper.