Jávorka Sándor - Soós Lajos (szerk.): A Magyar Természettudományi Múzeum évkönyve 29. (Budapest 1935)

Fejérváry, G. J.: Further contributions to a monograph of the Megalanidae and fossil Varanidae - with notes on recent Varanians

O. pannonïcus IVORM, and O. oentralis L. — it being in the latter species extremely developed, so as to form quasi a second proc. spinosus — occurs in many Reptiles, among them in Yaranians too,** but this formation, effectively belonging to the neural spine recess, is absolutely different from what we are discussing in Me­galania. the vertebrae of Megalania are. without being in any way pachyostotic, so heavy-built, so massive — rather hyperostotic — that it is quite natural that the formation of the orimental zygo­sphenes, as well as that of the zygantra depending upon them, shows quite different particulars than in the cases of light-built, slender skeletons. The ..irregular area above the neural canal on the anterior side" proves, however, both in position and structural details, a ty­pical zygosphene, and my interpretation concerned has been strengthened upon the autoptical evidence offered by the splendid vertebra our Museum owes to the much obliging generosity of Mr. LONGMAN. Besides, my ^identification has been smoothly accepted, without the slightest objection, by a series of specialists, so by Prof. BROILI. Dr. CAMP and Baron NOPCSA, who all. since, refer to Me­galania as possessing vertebrae provided with zygosphenes and zy­gantra. In the characterization of the group referred to by Baron NOPCSA. 58 under the name of „Megalaninae" this author very cor­rectly remarks: .,small zygosphen present", in opposition to his Do­lichosaurinae — gathered with the former ..subfamily" and with his ..Varaninae" into the compass of his (nomenclaturally non-valid) fa­mily ..Platynotidae" — which „a re stated to possess a .,strong zy­gosphen". By this morphologically exact statement my previous mistake has been silently emended in literature, and, so Far as the actual estimation of the detail in question be concerned, nothing re­mains to be added to the subject. Closely connected with considerations bearing upon the biono­mical and biohistorical evaluation of the structure peculiar to Mega­lanian vertebrae, One point is still left to be here discussed. Ihat point consists in the determination of the systematical position, i. e. the phylogenetical relation, of this Australian Reptile. In my Mono­graph of the fossil Varanidae and the Megalanidae (1916) I assigned the systematical place of the latter family, provisionally, within the bounds of the Suborder Platynota I). & B. (lam.) emend. TTÍRBR. (sensu IÜRBR.). marking the conditionality of such arrangement by 57 Cfr. LONGMAN, op. cit. p. 22. 58 The Genera of Reptiles. Palaeobiologica, 1. Band. 1928, p. 177.

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