L. Forró - É. Murai szerk.: Miscellanea Zoologica Hungarica 6. 1991 (Budapest, 1991)

Nesemann, H.: Zoogeography and composition of leech fauna of Danubian lowland rivers in the Kisalföld compared with some molluscs (Hirudinea, Gastropoda)

1927, SAPKAREV 1970). There it is a substitute for E. nigricollis. The leech E. testacea is very rare in the hilly and mountainous areas of Central Europe, where its range is restricted. The species has not been found anywhere in the upper reach of the River Danube yet. The northwestern distribution in this river basin was confirmed in Austria. The flood plain of the lower River March represents the border of the settled area. Dina punctata Johansson, 1927 (Fig. 8) Localities: Danube at Érd-Ófalu (H), km 1625.4, 30. Jan. 1991, leg. WIMMER & NESEMANN; Danube at Budapest, Szabadság-híd (H), km 1645.3, 15. Oct. 1988; Danube at Szob (H), km 1706.6, Oct. 1989, leg. PUKY & NESEMANN; Danube at Vác (H), km 1680.4, Oct. 1989, leg. PUKY & NESEMANN; Danube at Nagymaros (H), km 1694.7, Oct. 1989, leg. PUKY & NESEMANN; Danube near Gönyü (H), km 1791.4, 23. Oct. 1990; Mosoni Duna at Dunaszeg (H), 23. Oct. 1990; very common in the Danube (A) between Wolfsthal (km 1876) and Stockerau (km 1950), see NESEMANN (1989a); March (Morava) near Markthof (A), 24. Mar. 1990; Lajta at Mosonmagyaróvár (H), 3. Apr. 1990; Leitha at Nickelsdorf (A), 26. Oct. 1990; Leitha at Hollern (A), 26. Oct. 1990; Leitha near Pachfurth (A), 27. May 1990; Leitha at Bruck (A), 3. Jun. 1990; very common in the middle reach of the River Leitha (A) between Bruck and Ebenfurth (19881990). The closely separated species seems to belong to a monophyletic group of the subfamily Trochetinae, including D. apathyi, D. stschegolewi (LUKIN & EPSHTEIN 1960) and several undiscussed forms. D. punctata is widely distributed in the Ebro-, Rhine-, Po- and upper Danube Basins. It originated from the western Paleoeuropean or Euromediterranean fauna (KINZELBACH 1987), unlike D. apathyi and D. stschegolewi, which originated from the eastern (Pontosarmatian) fauna. D. punctata is a characteristic member of the biocoenosis in fast flowing mountain streams and circum-alpine rivers with a strong bed load transport of round stones. The groundtype of distribution pattern is similar to that of rheophilic Cyprinid fish Leuciscus souffia (Risso, 1826). D. punctata is mainly located at altitudes higher than 300 meters above sea level. The species is the most abundant leech in the upper Danubian Basin in Germany and Austria. It becomes rare in the Hungarian Danube around Budapest. Its occurrence spreads downstream to Érd. Here D. punctata lives sporadically at undercut slopes on gravely and stony bottoms. In the following stretches D. lineata gradually becomes dominant compared to the other species listed (Szalkszentmárton, Dunaföldvár). In general Trocheta (ylindrica (March, Thaya), Dina punctata and D. lineata (Danube, Lajta) occur mainly allopatrically. Dina apathyi Gedroyc, 1916 (Fig. 1) Localities: Ráckevei (Soroksári) Duna near Dunaharaszti (H), 20. Jan. 1989, leg. FORRÓ & NESEMANN, 9. Oct. 1989, leg. PUKY & NESEMANN. It is a Ponto-Caspian species which is rarely found, but widely spread in the drainage area of the Black Sea and Caspian Sea. A general description, based on the Hungarian records, was previously given (NESEMANN 1991b). This semiaquatic species inhabits lowland rivers and lakes. It prefers stagnant water bodies of secondary branches, wherein it has already been reported by MIKUSKA & GEC (1970).