Dr. Éva Murai szerk.: Miscellanea Zoologica Hungarica 2. 1984 (Budapest, 1984)

Varga, A.: The Cochlostoma genus (Gastropoda, Prosobranchiata) in Yugoslavia. I. Anatomical studies

Cochlostoma septemspirale riedeli Varga 1980 Material: Montenegro, 20 kms from Ribarice towards Rozaj, on rocks along the road, coll. L. Pintér, P. Subai and A. Szigethy, 21. 7. 1972. Paratype (Hungarian Natural History Museum, Budapest, Figs 25-29). Female reproductive system: unknown. Male reproductive system (Figs 25-29). Thin sperm duct opening at apex of prostrate with an opening slightly shifted to side. Distal part of prostrate definitely disjunct from proximal part which, in general, regularly and strongly subdivided. Terminal part of prostrate, contrary to C. s. heydenianum and bosniense

, continuening as a thin tube and with a narrow opening into mantle cavity at rectum at posterior end of animal. Prostrate connected to penis through a thin and relatively long closed tube. Penis thin and long, finely rugose, tapering. Vas deferens undulate inside penis. Length-width ratios 8.2:1 and 8:1 (Fig.29). In young speciments this ratio 4.7:1 (Fig. 28). Remarks on penis development as based on Figs 27 and 28: intensive growth of penis in same period as in heydenianum and bosniense. Aperture of illustrated specimen translucent, no thickenings observed. Prostrate developing earlier than penis (Fig. 27). Contrary to heydenianum, mantle margin simple, not protruded. Cochlostoma scalarinum scalarinum (A. & J. B. Villa 1841) Material: Hrvatska, Starigrad, Isle of Hvar, coll. L.Drimmer, 12. 8. 1983 (Figs 30-38). Female reproductive system (Fig, 30): Oviduct thin and sinuous. SBC joining oviduct at about apex of uterus gland. Seminal receptacle positioned twice as far from this point as length of SBC. Seminal receptacle clearly distinct from oviduct, with a length-with ratio of 2:1. Sac of bursa copulatrix elongated, with A:B ratio of 1:2. 5. Uterus gland strongly developed, opening into mantle cavity close to rectum. Male reproductive system (Figs 30-38): Typical form possessing macropenis. Prostrate evenly thickened (Fig. 32), narrowly opening into mantle cavity near rectum at posterior end. Prostrate connected to penis with a short tube. Free section of penis strongly elongated and pointed. Length-width ratio on free part ranging from 5.6:1 to 6. 5:1 (Figs. 31, 33, 34). Vas deferens undulate inside penis. Micropenis form (Fig. 3 5) with short, thick, curved and abruptly tapering penis with a length-width ratio of 3.1:1. Cochlostoma scalarinum hirci (Hire 1881) Material: Hrvatska, Velebit Mts, Jablanac, on limestone rocks over a sea inlet (Nature Reserve), Ca, extremely dry habitat, coll. A. Wiktor, 25-26. 5. 1982 (Figs 39-47). Female reproductive system (Fig. 39): Oviduct thin and undulate. SBC joining oviduct at about apex of uterus gland. Seminal receptacle positioned from this point at a distance 1.1 times longer than SBC length. Seminal receptacle clearly separated from oviduct, lengthwidth ratio about 8:1. Sac of bursa copulatrix elongated, A:B ratio is 1.2:1. Uterus gland as in nominal form. Male reproductive system (Figs 40-47): Contrary to nomine' form, no clear distinction between two penile forms "observed in the sample. The size differences among specimens illustrated are due to fluctuations in the population or to the contraction of penis. Prostrate (Fig. 4) as in nominal form, length-width ratios being 5. 2:1, 5.3:1, 5:1, 4:1, 5.1:1, 5.1:1, respectively (Figs 40, 42-47). Vas deferens linear or undulate inside penis. The so-called micropenis form was not encountered in the material examined. Cochlostoma elegáns elegáns (Clessin 1879) Material: Hrvatska, Velebit Mts, above Karlobag in a dry gorge, elmwood, Ca, coll. A. Wiktor, 27.5.1982 (Figs 48-51). Sexes differing in colour. Shells of females uniformly bright. Shells of males with bright ground colour and upper whorls dark violet. Female reproductive system (Figs 48-49): Oviduct thin and undulate. SBC joining oviduct below apex of uterus gland. Seminal receptacle positioned from this point at a distance 1. 5 times longer than SBC length. Seminal receptacle a merely slightly inflated part of oviduct. Sac of bursa copulatrix one-sided. A:B ratio 1:4. Uterus gland fully developed over its total length, opening into mantle cavity at rectum. The specimen illustrated in Fig. 49 has a fully developed shell but the reproductive system is deformed.