Szabó János szerk.: Fragmenta Mineralogica Et Palaentologica 26. 2008. (Budapest, 2008)

Genus Epulotrochus COSSMANN, 1918 Type specie s: Trochus epulus D'ORBIGNY, 1853 According to FISCHER & WEBER (1997), Trochus epulus D'ORBIGNY, 1853 can not be regarded as an independent species, it would be a morphotype within Trochus acteon D'ORBIGNY, 1853. Consequendy, this latter species should be regarded as the type species of genus Epulotrochus COSSMANN, 1918. However, in the available material Epulotrochus acteon is doubtlessly found, but the "epulus" morphotype is unidentifiable, disregarding some uncertain cases in the "background" collection. Most of the simple conical forms belong to different species and specimens of transitional shape are also absent, in spite of the rather high frequency of Epulotrochus in the Hierlatz Limestone fauna. These facts suggest that unification of the two species was most probably unjustified, therefore both species are treated here as existing ones. Epulotrochus COSSMANN, 1918 is established for anomphalous shells of conical outline, consisting of flush whorls and simple peristome. Studying genera of the subfamily, MONARI et al. (1996) recognised that the firmest basis to distingush genera of the Proconulinae are the peristome, mainly the inner lip characters; the ornament can provide secondary tools. In accordance with this experience, here the original interpretation of Epulotrochus will be modified. In the Hierlatz Limestone materials, some ornamented shells of Epulotrochus shape and peristome morphology are found, they will be added to the genus. Figure 60 — Epulotrochus acteon (D'ORBIGNY, 1852) (A-F) and Epulotrochus tuherculatus n. sp. (G-P) from the boxes labelled as "Trochus epulus D'ORBIGNY" in STOLICZKA's (1861) collections (true Epulotrochus epulus not found). — 11 a—d: copv of Trochus epulus figures from STOLICZKA (1861) Tafel I; A-F: one of the Epulotrochus acteon (D'ORBIONY, 1852) specimens (GBa 2008/69/10/1/1) in apertural (A, D), dorsal (B, E) and basal (C, F) views, A-C = xl, D-F = x2; G-L: the paratype of Epulotrochus tuberculatns n. sp., apertural (G, J), dorsal (H, K) and basal (I, L) views, G-I = xl, J—

L = x2; M—P: holotype of Epulotrochus tuherculatus n. sp. with well preserved tubercled-ribbed juvenile shell part; M-N = Xl, O-P = X5. Epulotrochus acteon (D'ORBIGNY, 1853) (Figure 60: A-F) 1853: 'Trochus acteon D'ORBIGNY — D'ORBIGNY, p. 253, pi. 306, figs 13-16. pars 1861 : Trochus epulus D'ORBIGNY — STOLICZKA, p. 167, pl. 1, figs 11 a, c, ?d. 1997: Epulotrochus acteon (D'ORBIGNY, 1853) — FlSCI 1ER & WEBF.R, p. 97, pi. 19, figs 10 a-b. non 1997: Epulotrochus acteon (D'ORBIGNY, 1853) — FISCHER & WEBER, p. 97, pi. 19, figs 11 a-b. Material — In STOLICZKA's (1861) "originals collection", specimens of three species are saved as "Trochus epulus D'ORBIGNY", most of them (6) corresponds to Trochus acteon D'ORBIGNY but three other specimens represent two different species; one is diagnosed below as a new one (Epulotrochus tuherculatus n. sp.). About fifty specimens are available (GBa) from the Hierlatz Alpe and three ones from the Bakony Mts (HGM).