Szabó János szerk.: Fragmenta Mineralogica Et Palaentologica 26. 2008. (Budapest, 2008)

flattened in direction parallel to axis, outer lip simple, prosocline and strongly prosocyrt between selenizone and adapical suture, parietal lip not visible on available specimens. Callosity, connected to columellar lip in Ptychomphalus, present on visible part of periumbilical shell wall; umbilicus itself not füled in completely as usual in this genus. Sculpture — Shell completely smooth, disregarding extremely fine growth lines, visible under magnification and two weak spiral lines, delimiting selenizone. Remarks — Because EUDES-DESLONGCHAMPS's (1849) syntypes has been lost, FISCHER & WEBER (1997) selected a neotype and a paraneotype for Ptychomphalus heliciformis EUDES-DESLONGCHAMPS, 1849 from the collection, served as basis to D'ORBIGNY's (1851-1860) monograph, in which the specimens were identified as "Pleurotomaria rotellaeformis DLINKER". This peculiar situation well demonstrates the distinction problems between these two species with use only literature data. A material-based revision of these two species is badly needed. FISCHER & WEBER (1997) suggested two characters to distinguish P. helicifomns from P. rotellaeformis; the spire angle that is narrower (115-125°) in the latter and wider in the former species (130-140°), and the "little difference" between the shape of the columellar calluses. Because the latter difference has remained unspecified, the spiral angles of the Flierlatz specimens (135-153°) is the only objective character that seems to support to choose the name "heliciformis". Distribution — Fontaine-Etoupefour, (Calvados, France), Upper Pliensbachian (Spinatum Zone); Luxembourg, Lower Liassic; Hallstatt, Hierlatz Alpe and Schafberg (Austria), Upper Sinemurian (Oxynotum Zone); West Sicily (Italy), Upper Pliensbachian. Family Lophospiridae WENZ, 1938 Genus Worthenia DE KÖNINCK, 1883 Type species: Turbo tabulates CONRAD, 1835 Worthenia superstes SZABÓ, 1980 (Figure 20) 1980: Worthenia? superstes sp. n. — SZABÓ, p. 56, pl. 1, figs 10-1 1. 1991: Worthenia superstes S/.ABÓ, 1980 — INESTA et al., p. 20, pl. 1:1. Material — Five more or less fragmentary inner mould specimens with shell fragments (HGM, holotype J 9595). Measurements H HL HP D W AA AL holotype - 24.5 17 27 17 - 56° Shape — Gradate (pagodiform) shell, consisting of whorls with broad, slightly convex ramp, bordered by rather marked angulation at midwhorl. Selenizone carinalike and coincides with angulation; periphery also keel-like. Outer face slightly concave, and abapically feebly inclined toward axis. Base convex and anomphalous. No peristome part preserved in available material. Sculpture — Whorls covered by fine spiral lines, while base ornamented with stronger spiral cords. Selenizone crenulate and bearing also two or three spiral threads. Growth-lines distincdy prosocline, slighdy prosocyrt on ramp, prosocyrt on outer face and opisthocyrt on base. Remarks — The shape and the ornament of the selenizone distinguish this form from the similarly gradate Bathrotomaria and Sisenna species. Its ornament can be reconstructed from the shell remnants (arrows in Figure 20: C). In the first description, the genus name seemed questionable but the subsequent, extended studies on Jurassic and Triassic gastropods confirmed its tightness. This species belongs to a survivor line of Worthenia. After long Palaeozoic—Triassic history

7 and species richness, this genus became really extinct most probably during the Toarcian crisis. Distribution — Lókút, Kericser (Bakony Mts), beds with Obtusum to Ibex Zone mixed fauna, and Stokesi Zone (Upper Sinemurian? to Upper Pliensbachian); Mola Hill (Novelda, Alicante, Spain), Pliensbachian. Figure 20 — Worthenia superstes SZABÓ, 1980; refiguration of the holotype —A-B: "apertural" and dorsal views, xl; C: magnified dorsal view to show remnants of the shell with ornament (arrows), x3.