Szabó János szerk.: Fragmenta Mineralogica Et Palaentologica 26. 2008. (Budapest, 2008)

However, these characters are species distinctive therefore the necessary corrections are given below now; the Bakony Mts (Úrkút) specimen is distinguished as Asterohelix urkutensis n. sp. Because the revision of STOLICZKA's (1861) originals has verified that Discohelix spinicosta STOLICZKA, 1861 also belong to Asterohelix, there is no reason to change the type species in accordance with the ICZN (1999, Article 70.3). It means at the same time, that the figures in the original designation of Asterohelix SZABÓ, 1984 do not refer the type species; its true images are shown here (Figure 13: A-E). Three Asterohelix species from the Mediterranean Jurassic have the typical "asterisk" outline at least in juvenile stage and the other morphological details show also their close phylogenetical connections [Asterohelix urkutensis n. sp., Lower(?) Sinemurian; Asterohelix spinicosta (STOLICZKA, 1861), Late Sinemurian; and Asterohelix mariae (M. GEMMELLARO, 1911), Late Pliensbachian)]. Recently FISCHER (2001) and GRÜNDEL (2003a, 2003b) identified further, more or less different species as Asterohelix from the Arabian Peninsula and stable Europe, respectively. Asterohelix spinicosta (STOLICZKA, 1861) (Figure 13) 1861: Discohelix spinicosta STOL. — STOLICZKA, p. 185, pi. 3, fig. 15. non 1984: Asterohelix spinicosta (STOLICZKA, 1861) — SZABÓ, p. 68, fig. 2. Lectotype — GBa 2008/69/36/1 Material— GBa 2008/69/36/1-8 in the "originals collection". Measurements H HL HP D W AA AL lectotype *3 *2.8 *2.3 9 2.5 *175 159° Figure 13 — Asterohelix spinicosta (STOLICZKA, 1861) — 15 a-c: copy of the original figures (from STOLICZKA 1861, Tafel III); A-B: lectotype in apical (A) and outer side (B) views, x2; C: apical view of a paralectotype, best showing the ornament and shape of the adult last whorl, X2.5; D: imprint of a paralectotype, best showing the basal sculpture, x3; E: spiral view of a paralectotype (juvenile shell), X4. Shape — Strongly flattened shell of feebly trochospiral coiling with cyrtoconoidal spire outline. Early spire part planispiral, but trochospiral coiling develops from third whorl. Protoconch itself poorly preserved in all specimens but visible parts suggest similar morphology to that of Asterohelix urkutensis n. sp. (see below). No part of umbilical side of youngest whorls known. Whorls in umbilicus gradate, and more flattened than on spire side; outer side feebly convex and oblique to axis its adapical edge corresponds to periphery. No undamaged peristome was found; impression on inner mould shows its trumpet-like form with outward tapering lips (see Figure 13: A). Sculpture — From third whorl, carina with sparse spinulae develop along spire side suture, being fine incision initially but gradually becoming rather deeplv impressed while spinulae strengthen into prosocyrt ribs. Ribs periodically repeating, their spines become parabolic with growth on spire side (umbilical ones not observable). Keels of outer angulations gradually weaken on penultimate whorl and lacking from adult last whorl. From second half of penultimate whorl, parabolic nature of ribs vanishes, their strength and regularity decrease, but density and length increase. Ribs become suture to suture/angulation in length, and subregularly repeating collabral ridges in shape (Figure 13: A, C). Similar, but somewhat weaker and two times denser ribs developed in abaxial belt of umbilical side of last whorls. On same side, dense, short riblets terminate in nodulae, formed along adaxial angulation (Figure 13: D). Growth-lines fine, hardly visible on early whorls then gradually strengthen into dense "growth-riblets" that run from spiral suture to umbilical suture across outer side. "Growth-riblets" became truncated at parabolic ribs. No trace of spiral ornament found. Remarks — The feebly trochospiral shells and the last whorl morphology, which is significantly different from that of the earlier whorls, distinguish Asterohelix spinicosta from Asterohelix urkutensis n. sp. (see below). Asterohelix mariae (M. GEMMELLARO, 1911), is distinguishable by its last whorl, having neither carinae nor spines, and by its concave spire. Distribution — Hallstatt, Hierlatz Alpe: Hierlatz Limestone; Sinemurian (Oxynotum Zone).