Szabó János szerk.: Fragmenta Mineralogica Et Palaentologica 24-25. 2007. (Budapest, 2007)

4 Ol LM, A. Genus Argyrotheca DALL, 1900 Argyrotheca cuneata (Risso, 1826) (Figure 3: 1-7) 1990: Argyrotheca cuneata (RlSSO, 1826) — BlTNER, pp. 138-140, text-figs 5-6, pi. 4, figs 1-9. (cum. syn.) 1993: Argyrotheca cuneata (RlSSO, 1826) — BlTNER, pp. 149-150, pi. 2, figs 1-6, pi. 3, figs 1-6. 2000: Argyrotheca cuneata (RlSSO, 1826) — BlTNER & PlSERA, p. 9, pi. 1, figs 1-7. 2002: Argyrotheca cuneata (RlSSO, 1826) — TOSCAXO& SORGENTE, pi. 20, figs 2(?)-3. 2002: Argyrotheca cf. cuneata (RlSSO, 1826) — K< AYALEWSKI et al., fig. 2A. 2003: Argyrotheca cuneata (RlSSO, 1826) — KROH, p. 148, pl. 1, figs 1-3. 2004: Argyrotheca cf. cuneata (RlSSO, 1826) — SlMÔES et al., pp. 523-524, text-figs 3A-B, text-figs 5F-N. 2004: Argyrotheca cuneata (RlSSO, 1826) — BlTNER & KAIM, p. 196, figs 2A-C. 2004: Argyrotheca cuneata (RlSSO, 1826) — BlTNER & DULAI, p. 78, pl. Ill, figs 9-10. Material — 147 specimens (94 complete shells, 25 dorsal valves, 28 ventral valves) from Bánd. Remarks — Although the genus Argyrotheca was widely distributed within the Central Paratethys, A. cuneata was less frequent than A. cordata. The ornamentation and the shell outline of A. cuneata may be similar to Megathiris detruncata. However, it is much smaller, having fewer and not so strong ribs, and it has only one septum on the dorsal valve. The specimen figured by TOSCANO & SORGENTE (2002, fig. 2) shows very strong ribs, but without informations on the internal characters it is impossible to decide between A. cuneata and M. detruncata. A. cuneata can also be distinguished from A. cordata by having broad ribs on the external surface of both valves, and by the internal characters (no tubercles at the internal anterior margin of the valves). Some earlier authors distinguished the Tertian' specimens under the name costulata, and suggested that costulata could be the Miocene ancestor of the Recent cuneata. BlTNER (1990) synonymized these two species and she also discussed 1990 Argyrotheca cordata (RlSSO, 1826) 1993 Argyrotheca cordata (RlSSO, 1826) 2000 Argyrotheca cordata (RlSSO, 1826) 2002 Argyrotheca cordata (RlSSO, 1826) 2004 Argyrotheca cordata (RlSSO, 1826) 2004 Argyrotheca cordata (RlSSO, 1826) Material — 275 specimens (191 complete shells, 39 dorsal valves, 45 ventral valves) from Bánd, 2 specimens (2 dorsal valves) from Devecser. Remarks — This is the most frequent species within the studied material. In some earlier papers it was reported under the name A. neapolitana (SCACCHI) but the name cordata has priority over the name neapolitana (LOGAN, 1977). The two BOETTGLR's (1901) species, A. subcordata and A. subcuneata are also conspecific with A. cordata, because they are within the intraspecific variability ranges of this species (BlTNER 1990). BlTNER (1990) has also synonymized DEMoRGAN's (1915) species (Cistella laevigata, C. Marie), as well as FRIEDEERG's (1921) species {Cistella dertomutiensis, C. ^boroviensis) with A. cordata. MEZNERICS (1943) used three different names for A. cordata specimens: neapolitana, subcordata, subcuneata. This species is easily distinguishable from A. cuneata by the internal features: the internal anterior margin of both valves is covered with tubercles (4—20, the similarity with species A. squamata (ElCHWALD) and A. jacksoni COOPER. Palaeoecology — Attached bv a short pedicle to firm substrates in cryptic habitats (under boulders, undersides of overhangings, coralligène, cave walls and roofs). Its depth range is similar to that of A. cordata, from few metres to more than 600m, but more frequent in shallow waters (infralittoral-circalittoral) (LOGAN 1979, 1983, 2003; LOGAN & NOBLE 1983; LOGAN et al. 2004). Distribution — Well-known species from the Miocene to Recent. Miocene: Poland (BlTNER 1990; POPIELBARCZYK & BARCXYK 1990; BlTNER & PlSERA 2000; BlTNER & KAIM 2004); Austria (KROH 2003); Hungary (MATYASOVSZKY 1880; BlTNER & DULAI 2004); Bulgaria (BlTNER 1993); Italy (DAVIDSON 1870; SACCO 1902); Recent: Mediterranean Sea (LOGAN 1979, 2003; LOGAN & NOBLE 1983; BRUNTON 1988; LOGAN et al. 2002, 2004); Atlantic Ocean (BRUNTON & CURRY 1979; LOGAN 1983, 1988, 1993; KOWALEWSKJ et al. 2002; SlMÔES et al. 2004). from small to large specimens). The triangular median septum shows 3-4 (BlTNER 1990) or 5-6 (LOGAN 1979) serrations. The Hungarian specimens have 4 serrations (Figure 4: 5, 7-9). Palaeoecology — The species is attached by a short pedicle to firm substrates in cryptic habitats (under boulders, undersides of overhangings, coralligène, cave walls and roofs). Its depth range is from few metres down to more than 600m, but it is more common in shallow waters (infralittoral-circalittoral). Densities of up to 100 specimens/m

2 were recorded (LOGAN & NOBLE 1983; LOGAN 2003; LOGAN et al. 2004). Distribution — Well-known species from the Miocene to Recent. Miocene: Poland (BARCXYK & POPIELBARCZYK 1977; STUDENCKI 1988; BlTNER 1990, 2002; POPIEL-BARCZYK & BARCXYK 1990; BlTNER & PlSERA 2000; BlTNER & KAIM 2004); Ukraine (FRIEDBERG 1921); Austria (DREGER 1889); Hungary (MATYASOVSZKY 1880; Argyrotheca cordata (RlSSO, 1826) (Figure 4: 1-9) — BlTNER, pp. 140-143, text-figs 7-8, pi. 5, figs 1-14, pi. 7

, fig. 1. (cum. syn.) — BITNER, p. 150, pi. 4, figs 1-5, pi. 5, figs 1-8. — BlTNER & PlSERA, pp. 9-10, pi. 2, figs 1-9. — TOSCAN o & SORGENTE, pi. 20, fig. 1. — BITNER & KAIM, pp. 196-197, figs 2D-K, 3A-E. — BlTNER&DULAI, p. 74, pi. Ill, figs 2-8.