Szabó János szerk.: Fragmenta Mineralogica Et Palaentologica 23. 2005. (Budapest, 2005)

16 HÍR Table 4 — Individual measurements and morphological data of Collimys dobosi

ri. sp. mi molars from Felsőtárkány 3/2. N° inv, MMP Length Width Morphology mm mm Figure 16 2003. 206. 1.78 1.20 CI 2003. 207. figure 1. 1.85 1.20 CI paratype 2003. 208. 1.92 1.23 CI 2003. 209. 1.96 1.23 CI 2003. 210. 1.89 1.20 Dl 2003. 211. figure 17: 6 1.82 1.09 CO 2003. 212. 1.82 1.19 CI 2003. 213. figure 17: 5 1.86 1.16 Bl 2003. 214. 1.82 1.18 Dl 2003. 215. 1.85 1.20 CO 2003.216 1.95 1.22 CI 2003. 217. 1.93 1.30 Bl 2003. 218. 1.96 1.20 Bl 2003. 219. figure 17:2 1.83 1.23 Dl 2003. 220. 1.86 1.20 AO 2003. 221. 1.85 1.26 B0 2003. 222. 1.18 CO 2003. 223. 1.89 :. :

1 Dl 2003. 224. figure 17: 1 1.89 1.18 AO 2003. 225. figure 17: 8 1.85 1.23 DO 2003. 226. 1.81 1.13 CI 2003. 227. figure 17: 4 1.81 1.18 B0 2003. 228. 1.82 1.16 2003. 229. 1.78 1.16 CO 2003. 230. 1.90 1.22 Bl 2003.231. 1.86 1.22 2003. 232. Dl 2003. 233. 1.79 1.12 CI 2003. 234. 1 .''5 1.25 2003. 235. 2.00 1.26 Dl 2003. 236. 1.75 1.19 Bl 2003. 237. 1.82 1.19 CO 2003. 238. 1.89 1.26 AO 2003. 239. 1.96 1.22 Bl 2003. 240. 1.96 1.15 CO 2003. 241. figure 17: 9 1.81 1.15 Dl 2003. 242. 1.79 1.18 Dl 2003. 243. AO 2003. 244. 1.89 1.30 2003. 245. 1.88 1.13 Dl 2003. 246. 1.96 1.26 CO 2003. 247. 1.83 ;.i

i

> Di 2003. 248. 1.85 1.16 C1 2003. 249. 1.99 1.20 CI 2003. 250. 1.83 1.16 AO 2003. 251. 1.93 1.15 Dl 2003. 252. 1.76 1.20 CI 2003. 253. 1.92 1.22 DO 2003. 254. 1.86 1.23 2003. 255. 1.85 1.16 AO 2003. 256. 1.86 1.23 B( i 2003. 257. 2.09 1.26 Bl 2003. 258. 1.92 1.16 CI 2003. 259. 1.79 1.18 Dl 2003. 260. 1.7 1 1.15 CO 2003. 261. 1.85 1.19 B0 2003. 262. 1.78 1.16 Bl 2003. 263. 1.99 1.19 CI 2003. 264. 1.82 1.13 Bl 2003. 265. 1.81 1.16 Al 2003. 266. 1.90 1.25 CI 2003. 267. figure 17: 3 1.96 1.16 Al 2003. 268. 1.89 1.22 2003. 269. 1.90 1.16 CI 2003. 270. 1.96 1.23 CI 2003. 271. 1.74 1.13 CI 2003. 272. 1.79 1.08 CO 2003. 273. 1.82 1.13 CO 2003. 274. 175 1.16 2003. 275. 1.75 1.06 CI 2003. 276. 1.71 1.1 6 Bl 2003. 277. 1.93 1.23 Bl 2003. 278. 1.75 1.19 2003. 279. 1.72 1.15 2003. 280. 1.82 1.13 CI 2003. 281. 1.81 1.12 Dl 2003. 282. 1.81 1.20 B0 2003. 283. 1.81 1.12 CO j\e 76 76 Min. 1.71 1.06 Max. 2.09 1.30 X 1.8575 1.18737 Standard deviation 0.06178 0.03679 Discussion Eleven localities containing Collimys finds are listed by KÄLIN & ENGESSER (2001): Steinheim, Petersbuch 31, Le Locle/Sous-le-Stand, Ottenberg 3, Chräzerentobel 655m, Petersbuch 10, Belchatow A, Nebelbergweg, Hammerschmiede, Dorn-Dürkheim, Eichkogel. Schernham was published as the type locality of Pseudocollimys DAXNERHÖCK, 2004. Adequate descriptions are published only from Steinheim, Nebelbergweg and Eichkogel. C. dobosi n. sp. shows a mosaic of archaic and progressive markers. All investigators of the Collimys remains emphasize the importance of the absence of the lingual anterolophule in the M

2 molars as a characteristic marker of the Collimys genus. This crown element is surprisingly frequent in the population of Felsőtárkány. It is probably an archaic marker of this sample. Another archaic morphological character is the frequent anterior metalophule in the crown of the M

3 molars. It is found in C transversus, but it is missing in C longidens. From chronological point of view C. dobosi is situated between C. transversus and C. longidens but it does not mean an intermediate evolutionär}' status. Beyond the above mentioned frequency of the lingual anterolophule of the M

2 molars the generally larger dental measurements, the relation of the Lm? and L1T13 and the differences in the structure of the anteroconid-anterolophulid region of the ml molars suggest that C. dobosi can not be the ancestor of C. longidens. The occurence of the additional enamel ridges in the metalophule of IVF-M

2 are progressive markers, because these are found in the tooth crowns of C. primus. Another important aspect of C. dobosi at Felsőtárkány 3/2 is its dominance in the rodent fauna. 54.5% of the identified intact rodent molars refer to Collimys. This fact is interresting, because the genus is known as a rare element of the faunas in Switzerland, Germany and Austria. Currendy the paleoecological significance of the high frequency of Collimys in the Felsőtárkány 3/2 fauna is not understood. The next most common rodent genus for Felsőtárkány 3/2 is Anomalomys (14%) (HÍR 2002c). The majorit}'' of the rodent assemblage is characterized by two species having tendency towards hypsodonty. In spite of this fact the paleoflora of Felsőtárkány is dominated by