Szabó János szerk.: Fragmenta Mineralogica Et Palaentologica 21. 2003. (Budapest, 2003)

Tardos, Kisgerecse — Twenty-one brachiopod species were collected at the northwestern slopes of the Kisgerecse (DUI-AI 1998b), from the basal layers of the Pisznice Limestone. The fauna is diverse but no ammonites were found. Rhynchonellids are prevailing (43%), terebratulids (36%) and spiriferinids (19%) are also frequent, whereas athyridids are rare (2%). The most frequent brachiopod taxa are Liospiriferina (19%), Phymatothyris (16%), Zeilleria (14%) and "PJynchonella" triquetra (13%). Calcirhynchia (9%) and Piarorlynchia (8%) are also relatively common but all the other genera are rare (Prionorlynchia, Cirpa, Cundrbynchia, Gibhirhynchia, Lobotlyris, Koninckodonta) (Figure 4). Tardos, Vöröshíd quarry

— At this locality the Pisznice Limestone unconformably overlays the Upper Triassic Dachstein Limestone. Bed-by-bed collection was done from an approximately 14 m thick sequence, which were clivided into two parts (DULAI 1998b). Twenty-six brachiopod species were distinguished at this locality. Ammonites were found only in the upper part, which are Early Sinemurian (Arnioceras semicostatum Zone, PÁLFY pers. com.). The lower part of the sequence probably belongs to the Arie tries bucklandi Zone (see DULAI 1998b). The stratigraphie distribution of the brachiopod species were compared to those of the Lábadan localities (Tölgyhát and Póckő) where no ammonites had been found (DULAI 1998a, b). The high number of brachiopods in common suggest that the studied strata at Tölgyhát and Póckő can probably be correlated with the lower, thick-bedded part of the Vöröshíd quarry. Thus the examined Lábatlan layers probably also belong to the Arterites bucklandi Zone. Rhynchonellids (45%) are prevailing the brachiopod fauna of this locality but terebratulids (37%) and spiriferinids (17%) are also common but athyridids are very rare (1%). The most frequent brachiopod genera are Linguithyris (22%), Cakirhynchia (18%) and Liospiriferina (16%). Beside them, Pisirlynchia (10%), Apringia (6%) and Phymatothyris (6%) are relatively also important. All the other genera are represented by only a few specimens {Prionorhynchia, Cirpa, Cuneirhynchia, Gibhirhynchia, "Rhynchonella, Dispiriferina, Labothyris, Zeilleria, Antiptychina, Rhapidothyris, Koninckodonta) (Figure 4). Figure 5 — Proportion of the brachiopod orders of the Hettangian localities in the Transdanubian Central Range. Depth control on taxonomic distribution and timing of intra-phylum niche replacement The bathymétrie and environmental distribution of the Late Triassic and Early Jurassic brachiopods were investigated by SANDY (1995a) in an evolutionary paleoecological study. SANDY found that the latest Triassic (Rhaetian) shallow water, high-energy associations are predominated by short-looped terebratulids, whereas athyridids are common components of deeper water, lowenergy environments. Spiriferinids, long-looped terebratulids and rhynchonellids occupy mid- to outer shelf environments. By the Early Jurassic (Pliensbachian) the shallow water brachiopod faunas are predominated by rhynchonellids. The short-looped terebratulids are common in relatively deeper shelf waters, whereas spiriferinids and long-looped terebratulids are abundant in basinal deepwater environments. These data were interpreted by SANDY (1995a) that an intra-phylum niche replacement took place within the Brachiopoda after the end-Triassic mass extinction. As a result, athyridids were replaced by spiriferinids and long-looped terebratulids in deep-water environments. Rhynchonellids appear to have diversified