Vörös A. szerk.: Fragmenta Mineralogica Et Palaentologica 12. 1985. (Budapest, 1985)

Holotype: Right mandible fragment with adult M-^-Mg teeth deposited in the Paleontological Department of the Hungarian Natural History Museum, Budapest; inventory no.: V. 84. 135.

(Pl.I, Fig.l). Paratypes

: 1. Bilateral maxillary fragment with M

1-M

3

, and M

2-M

3 teeth, respectively (V. 84. 136.), Pl. Ill, Fig. 1. 2. Bilateral maxillary fragment with M^M

3 and M

1-M

2 teeth, respectively (V. 84. 137), Pl. IV, Figs 3-4. 3. Left mandible fragment with Mj-Mg tooth-row (V. 84. 138), Pl. I, Fig. 2. 4. Right mandible fragment with I and M_-M tooth-row and with the fragment of the processus (V. 84. 139), Pl. I, Fig. 4 and Pl. ÍI, Fig. 4. Diagnosis: Spalacinae with medium-sized and slightly hypselodont teeth. Form of Jaw like that of Anomalomys

. On the lower incisor a longitudinal crest may be found. Its tooth-structure still preserves the complicated Anomalomys pattern, the line of folds and their wide angular arch are similar to those of the Prospalax

. The majority of the tooth-elements (94-95 %) constitute a continuous fold system. Stratum et locus typicus: The same as in the case of Anomalospalax gen.n. Other finds

: 9 mandible fragments with Mj-Mg tooth row, 2 maxillary fragment with M^-M

3 tooth row; total number of teeth: 37 M«, 25 M

2

, 38 Mg, 24 M , 25 M

z

, 19 M

d

. Description: Mandible: The corpus mandibulae is arched and with parallel edges. The processus coronoideus rises at the meeting point of M

2

~Mg, and in lateral view it covers Mg. The structure of the rhamus is the same as that of Anomalomys (VIRET & SCHAUB 1946, Fig. 2.). The tooth row inclines from the mandible incisor axis by 10-12°. The foramen mandibulae is located under the distal part of M,. M-: Holotype: It consists of continuous folds. In the medial fold (protoconid), which demonstrates the most Anomalomys features, a re-entrant can be found. Other finds

: Medium-sized, slightly hypselodont teeth. Its fold pattern is continuous. Lingually one, labially two re-entrants can be found. The anterior fold is of a drop shape and continuously transforms into a medial fold. There is a small bridge between the medial and the posterior folds. The enamel layer on the posterior fold is parallel. The fold pattern lingually is generally angularly arched. The inner part of the reentrants is slightly hollow. In case of the older abraded specimens, similarly as in case of the posterior lingualis, an enamel island develops because of the closing of the anterior lingual reentrant fold. In case of the abraded teeth the size and width increases, and the angularity of the fold arches is greater. In 28% of the Mj finds at Tardosbánya there are tooth element variations. In 23% of the Anomalomys remains the re-entrants sink into the medial fold, while in 5% of the cases the anterior fold is isolated from the other elements. M,: Holotype: The structure of the teeth is continuous. The Anomalomys re-entrant present in Mj is missing. Other finds

: The only difference from Mj is that the edge of the anterior fold, due to the adjoining surfaces of M

1-M

2

, is straight and not arched. In 35% of the M2 there are tooth element variations, 6% of which are of the Anomalomys type. It means that the enamel line of the re-entrants is not parallel and is irregulary crisped. The anterior fold is never isolated as in case of Mj, the isolation of the posterior fold is however 17%. M

3

: Holotype: The Z-shaped fold system is the same as in Mj and M2, but backwards it becomes thinner. The lingual end of the posterior fold (hypoconid) is isolated forming a separate small conid. Other finds

: The fold enamel line is uniform and parallel, and is sharply and angularly broken. In 42% of the Mg there are tooth variations. The complicated line of the anterior lingual re-entrant (7.5%) can be considered as an Anomalomys pattern. Two and half percent of the anterior folds and 7.5% of the posterior folds are isolated. The conid formation that can be seen on the holotype specimen is also present in 25% of the cases. Summing up the morphological patterns of the lower tooth-row, it can be stated that the Ano malomys patterns are the most frequent in Mj (23%), then in M

2 (21%) and are the less frequent in Mg (7.5%). Significant is the fact that the presence of this conservative feature is not consequent within any given specimen but appears rapsodically. This proves that Anomalospalax gen. n. is a transitional stage between the more complicated Anomalomys and the younger, simpler Spalacinae (? Prospalax)

. All these show that among the lower teeth the Mj and M„ are the most conservative. Fig. 1. Scatter diagrams of Anomalospalax tardosi (continuous) and Anomalospalax viretschaubi (dotted) from Tardosbánya and Polgárdi 4. The vertical axis is the lenght of tooth, the horizontal is the width in mm