O. Merkl szerk.: Folia Entomologica Hungarica 67. (Budapest, 2006)

Biology - Only the sexual generation is known. Adult wasps emerge from mid-May until mid-June. In the northern provinces development is somewhat later than in Lorestan, Phars and Ham provinces. The gall develops on both very young Fig. 129. Map of Iran, showing mountain chains and oak distributions (b = Q. brantii, c = Q. castaneifolii, i = Q. infectoria; 1 = Q. libani, m = Q. macranthera, p = Q. petraea)