S. Mahunka szerk.: Folia Entomologica Hungarica 58. (Budapest, 1997)

We also collected three species, which have had only one locality in the Great Hungarian Plain (Szél 1996). These three species were caught in the Ukrainian side: an abundant population of Carabus arcensis carpathus Born, 1902 (the dominant in the given habitat), Molops piceus (Panzer, 1793) (subdominant in the habitat) and Blethisa multipunctata (Linnaeus, 1758). Carabus arcensis carpathus Born, 1902 has more abundant populations in the Zempléni-hegység and the Aggteleki-karszt while specimens are known from the Mátra Mts. as well. Molops piceus (Panzer, 1793) is known from the Alpokalja, the Transdanubian Mountains and Northern Mountains as well as at Bátorliget. Blethisa multipunctata (Linnaeus, 1758) is rare in Hungary. Only one locality is known (Kalocsa) from the Great Hungarian Plain. Several species which are sporadic and (or) rare in the Great Hungarian Plain (Szél 1996) were also collected, from Hungary: Pterostichus melas (Creutzer, 1799) and from the Ukraine: Harpalus roubali Schauberger, 1928, Pterostichus latoricaensis Pulpan, 1965, P. melas (Creutzer, 1799) and Platynus livens (Gyllenhal, 1810). We detected both from the Hungarian and the Ukrainian side Carabus hampei Küster, 1846 which belongs to the subgenus Morphocarabus Géhin, 1876. This species has a Dacian distribution. About 12 subspecies of Carabus hampei Küster, 1846 are known (Lie 1992) from which C. hampei ormayi Reitter, 1896 lives on the Beregi-síkság and in Hungary only few localities are known [Nagyar, Szatmárcseke, Tarpa (Szél 1987) and Barabás: Kaszonyi-hegy (hill) (Szilágyi and Szálai unpublished datum)]. The locality of this subspecies detected in the Ukraine is the most remote north-westerly one. From the point of view of nature conservation it is important that this population lives in a very small, isolated forest patch. Therefore, the protection of the habitat is crucial to the survival of this population. The observed carabid species also prove that the fauna of the Beregi-síkság is biogeographically quite unique (Varga 1995). On the one hand, Carpathian effects can be experienced which are demonstrated in the occurrence of several mountainous species [e.g., Carabus arcensis carpathus Born, 1902, C. intricatus Linnaeus, 1761, C. zawadskyi Kraatz, 1854, Cychrus caraboides (Linnaeus, 1758), Molops piceus (Panzer, 1793) and Abax parallelus (Duftschmid, 1812)]. On the other hand, Dacian influence can be detected which is reflected in the presence of populations (e.g., Carabus hampei ormayi Reitter, 1896) heading from Transylvania to the west. These species often reach the most westerly border of their area in isolated populations. As the marginal populations are very sensitive, the extended protection of these local populations and their habitats are essential. We studied the biogeographical relation of the carabid fauna of the Beregi-síkság with the Caipathians in an indirect way by using multivariate statistical methods. The biotic contact of the Caipathians with the Aggteleki-karszt and the Zempléni-hegység (Varga 1995) can be proved with the occurrence of the same mountainous [e.g., Carabus glabratus Paykull, 1790, C. intricatus Linnaeus, 1761, C. zawadskyi Kraatz, 1854, C. variolosus Fabricius, 1787, Cychrus caraboides (Linnaeus, 1758), Molops piceus (Panzer, 1793)] and Carpathian [Carabus arcensis Herbst, 1784, Carabus obsoletus Sturm, 1815, 7)7chotichnus laevicollis (Duftschmid, 1812), Abax schueppeli Palliardi, 1825] carabid species (e.g., Móczár 1956, Papp 1958, Lie 1992, 1994). Thus, the Carpathian relation of the carabid fauna of the Beregi-síkság can also be demonstrated indirectly by comparing it with the carabid data of the Aggteleki-karszt and that of the Zempléni-hegység.