S. Mahunka szerk.: Folia Entomologica Hungarica 54. (Budapest, 1993)

Table 2. Habitat fidelity and habitat selection index of more abundant carabid species (represented by >100 individuals) collected by pitfall traps from six habitats of Nagy-szénás. (For the abbreviations, see chapter of Material and Methods) Species Habitat fidelity (HF) Habitat selection index(HSI) Species BR FO QC MF PN FA Habitat selection index(HSI) Leistus rufomarginalus (Duftschmid, 1812) -0.94 0.18 -0.18 0.29 0.14 0.02 1.75 Pterostichus melas (Creutzer, 1799) -0.15 -0.79 -0.01 0.10 -0.87 0.64 2.56 Pterostichus oblongopunctatus (Fabricius, 1787) -0.94 0.17 0.49 0.42 -0.80 -0.56 3.38 Abax parallelepipedus (Pilleret Mitterpacher, 1783) -0.85 -0.21 0.22 0.62 -0.36 -0.35 2.61 Molops piceus (Panzer, 1793) -0.84 0.16 0.00 0.63 -0.92 -0.17 2.72 Platyderes rufus (Duftschmid, 1812) -0.79 0.36 -0.19 0.58 -0.93 -0.19 3.05 Aptinus bombarda (Illiger, 1800) -0.99 0.88 -0.36 -0.28 -0.98 -0.51 4.00 The formation of the HF values within the main categories of the woodland associations depends on many factors. In the investigations of Niemelä et al. (1992) the soil moisture was the most important factor. Hurka (1986) proposes that the habitat and its microclimatic condition are the limiting factors for the development of carabids. Paje and Mossakowski (1984) found the pH-preference as a potential limiting factor. The availability of the prey is also an important factor (Loreau 1983). The HSI spreads between 1.75 and 4.00, and remains considerably under the maximum 6.00. This means, that is no outstanding habitat specialist carabid, but the „BR" is - with exeption of Pterostichus melas - not a favourable habitat for any species on the basis of the HF values. Leistus rufomarginatus has a low habitat selectivity. Besides „BR" this species is almost non-selective, which means that it is present with a homogeneous distribution among the forest habitats. Moreover, it seems that „PN" is optimal for this species only, because the HF value of the local dominant species, Abax paral­lelepipedus, is only under zero, but in the case of L. rufomarginatus this number is positive. The HSI of the other species is higher, those dispersion is quite heterogeneous. It means, that Leistus rufomarginatus can be regarded as a forest generalist species, while the others (especially Aptinus bombarda) are more or less forest specialists. There is a similar correlation in the HF values between „PN", „BR" and the other four associations in our data, and in the investigations of Jennings et al. (1986), namely between the clearcut and other forest conditions. We regard the „PN" and „BR" habitats to be a clearcut area for the majority of species in Nagy-szénás. The composition of species of „CC" was quite different from the previous ones, and the spectrum of species was also much larger. The species caught here are mostly habitat specialist carabids, among them we can mention the followings: Notiophilus germinyi Fauvel, 1863, Olisthopus rotundatus (Paykull, 1790), O. Sturmi (Duftschmid, 1812), Harpalus cordatus (Duftschmid, 1812) and Cyminclis axillaris (Fabricius, 1794), which are faunistically interest­ing species, too. Calathus cinctus Motschulsky, 1850 was collected also in this habitat. Its status was recently cleared after cross-breeding experiments and analysis of morphological data (Aukema 1990 a, b).

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