S. Mahunka szerk.: Folia Entomologica Hungarica 49. (Budapest, 1988)
Description. Male (holotype considerably damaged, abdomen deformed, legs mostly absent). Carapace 0. 88 long, 0.65 wide, grey-brown, with vague, dark, radial stripes and darker margin. Chelicerae 0.33 long. Legs pale brown. Tml - 0. 19. Palp (Figs 33-35): Cymbium without proximal processes. Paracymbium L-shaped. Tegulum directed obliquely forward as a cone. Lamella characteristica complex. Embolus large, well-sclerotized. Abdomen almost black, with two longitudinal rows of large spots on hind half fusing into a transverse band. Female unknown. Diagnosis: The new species joins the key se rlingi-group just created by WUNDERLICH (1985) and seems to be particularly closely related to L. spasskyi Tanasevitch, 1986, but differs from it by the shape of the distal part of the lamella characteristica: it is deeply biramous in L. spasskyi , but uniramous, terminally rounded and provided with a spiniform process in L. pepticus sp. n. Remarks: The keyserlingigroup comprises at present four species close not only morphologically, but by their ecological preference as well: L. keyserlingi (Äusserer, 1867) (Europe), L. quadrimaculatus Kulczynski, 1898 (Slovakia, Lower Austria), L. spasskyi Tanasevitch, 1986 (USSR: E Ukraine, W Kazakhstan) and L. pepticus sp. n. (E Mongolia). All of them distinctly prefer xerie habitats of a steppe(-like) to (semi-)desert type and populate (quasi)zonal communities with a xerophytous vegetation, occurring there among plants and/or under stones, but not in humid "microoases" usually inhabited by Lepthyphantes in such landscapes. NEW RECORDS OF LEPTHYPHANTES IN THE FAR EAST L. abiscoensis Holm, 1945. Kamchatka: tiCär Milkovo; Sakhalin: Vitnitsa River. L. alacris (Blackwall, 1853). Sakhalin: near Okha, Novoalexandrovsk; Kamchatka: near Zhupanovo, Petropavlovsk-Kamchatsky . L. angu latus (O. Pickard-Cambridge, 1881). Sakhalin: near Okha, Vitnitsa River. This somewhat surprising discovery of a N European species in the Far East is certainly far from unique, as similar Euro-Far Eastern disjunctions sometimes do occur among spiders (s. ESKOV 1987), though seem to be quite seldom. The present record in the Far East is particularly interesting in two respects. First of all, it has altered and made more complicated the known vicariance pattern in the species complex angulatus- geminus - bipilis. Secondly, it allows to speculate about the group's genesis. Its range nowadays: L. angulatus inhabits N Europe from the British Isles to the Urals, except for the latter's tundra zone, i.e. Malo- and Bolshezemelskaya Tundra, where it is substituted by L. geminus Tanasevitch, 1982; L. geminus occurs in the N Urals, in Siberia eastward up to Yenisei; L. bipilis Kulczynski, 1885 is restricted to the Far East (Kamchatka, Cisokhotia). Hence, E Siberia is a terra incognita. The find of L. angulatus in Sakhalin permits to regard this species as an ancestor both for L. geminus and L. bipilis which could have had in the former times a N Eurasian distribution. During the Pleistocene, the range of L. angulatus might have lain more southerly than nowadays, but some of its Siberian populations could have become isolated in smaller réfugia and transformed since into separate species, L. geminus and L. b i- pilis. Since the later warming of the climate L. geminus might have colonized Siberia and even N Uralia, where the boreo-European L. angulatus , which could have survived during the Ice Age and recolonized Europe, did not let a further southerly expansion of L. geminus. On the other hand, L. angulatus seems to have failed to colonize Siberia east of the Urals apparently due to the. presence there of both L. geminus and L. bipilis , its vicariants, but has retained an isolated eastern outpost in Sakhalin (and the Maritime Prov., where it has not been found as yet? ). L. bipilis Kulczynski, 1885. Kamchatka: near Zhupanovo, Paratunka. L. karpinskii (O. Pickard-Cambridge, 1873). Sakhalin: near Okha. Figs 31-35. Lepthyphantes enormitus sp. n. (31. 32), 3 holotype, Lepthyphantes pepticus sp. n. (33-35), <3 holotype: 31= left palp; 32= embolic division; 33= right palp; 34, 35= lamella characteristica
