S. Mahunka szerk.: Folia Entomologica Hungarica 44/1. (Budapest, 1983)
1. N. bonellii group Nebria (Alpaeus) bonellii (Ad.) Carabus Bonellii Adams, 1817, Mém. Soc.Imp.Nat. Moskau, 5j 301 (type locality: Caucasus). Nebria Marschallii Dejean, 1826: 255 (type locality: "montagnes du Caucase"). Nebria Marschallii , Chaudoir, 1846: 108 Nebria Bonelli var. cechenoides Reitter, 1885: 353 (type locality: "Swanetien"). Nebria Bonelli var. ardua Starck, 1894, Wien.ent. Ztg, 13_: 11 (type locality: "montis Schugus, Cauc. occ"). Nebria Bonellii var. viridimetallica Pic, 1911, Échange, 2J7: 153 (type locality: "Caucase occidental"). Nebria (Alpaeus) bonelli + ssp. cechenoides + var. ardua : BÄNNINGER, 1925: 272 Nebria (Alpaeus) Bonellii , Csiki, 1927: 275 Nebria (Alpaeus) bonellii , Kirshenblat, 1932: 91 Nebria (Alpaeus) Bonellii , Banninger, 1949: 130 Nebria (Alpaeus) gaedlke i Jedliőka, 1965: 213 (type locality: Westkaukasus, Umgebung Dombai), B.yn . n . Widely distributed in the Caucasus from Majkop to Bakuriani (for detailed localities see KIRSHENBLAT, 1932), not recorded from Armenia. Very variable species in colour and proportions of body. Typical form with dark violet head and pronotum and greenish copper elytra. Specimens with greenish metallic lustre of the upper surface have been described as var. viridimetallica Pic. Very rare var. ardua Starck is characterised by entirely red appendages. Large specimens with extraordinary thickened head have been described as var. cechenoides Reitt. Lectotype, cf (HNHM) of this variation, designated here, with labels: "Mamch, VIII. 98", "7, Nebria Bonellii v. cechenoides Rtt" by handwriting of E. REITTER. Moreover, in the collection of HNHM have two specimens of this variation with label "Utsch-Dere, Caucasus, Starck". N_. gaedikei Jedl. is the manifest synonym of N. bonellii Ad. Nebria (Alpaeus) bonellii heinziana ssp.n., fig. 1 In 1978 large series was collected by W. HEINZ in NE Anatolia. The area of the new subspecies is situated far from the nominal form. There are only small differences in external characters from the nominal form. Colour more constantly and intensively violet, only sometimes with faint greenish lustre. Pronotum somewhat wider (tabl. 1). Aedeagus (figs 14, 18-20) larger and stouter, paramerae (figs 24, 25, 29, 30) longer. Holotype, cf: Anatolia or., Obere Waldzone bei Altiparmak, 1500-1900 m, 6-7.VIII. 1978, leg. W. HEINZ (H). Paratypes: 3 cf, the same label (ISU, HNHM); 2 cf , 2 g, NO Anatolien, Barhal-yayla SW Sarigöl, 1900-2000 m, 7.VIII. 1978, leg. W. HEINZ (ISU); 2 cf , 1 g, NO Anatolien, Karckal-Otingo, 2000-2800 m, Hochweidenzone, 11 .VIII. 1978, leg. W. HEINZ (ZIL, ISU); 2 cf, 1 g, NO Anatolien, pass siidl. Ikizdere, 1800-2500 m, 21.VIII. 1978, leg. W. HEINZ (ZIL, ISU). 2. N. fischeri group Nebria (Alpaeus) fischeri Fald., fig. 2 Nebria Fischeri Faldermann, 1835: 31, tabl. 3, fig. 2 (type locality not mentioned) Nebria Hyantis Reitter, 1899: 195 (type locality: "Tschoroch, Fluss südlich von Batum"), syn .n . Nebria (Alpaeus) fischeri , Banninger, 1925: 273 Nebria (alpaeus) Fischeri , Csiki, 1927: 378 Nebria (Alpaeus) Hyantis , Csiki, 1927: 381 Nebria (Alpaeus) hyantis . Kirshenblat, 1932: 91 Nebria (Alpaeus) fischeri , Kirshenblat, 1932: 92 Nebria (Alpaeus) hyantis . Banninger, 1949: 130
